CACTIBOTANY AND CULTIVATION
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Botanical terms and comparisons

Airampoa, Brasiliopuntia and Consolea · descriptions, former names and research
Leuenbergeria and Pereskia · species, former names and research
Ariocarpus · studies and flower comparison
Astrophytum research data

Comparisons from the articles gathered in one place. On a narrow screen, the table can be scrolled horizontally.

Names of cactus parts

Structural elements and how to recognize them
ElementMeaningHow to interpret it
ShootThe stem together with leaves and buds, even when leaves are highly reducedThe green body does not include roots. Shoot and stem are sometimes used as shorthand, but are not fully equivalent.
Growing tipThe region containing the shoot apical meristemNew sections of the axis and organ primordia form here. Not every areole is a growing tip.
AreoleA specialized short shoot with its own meristematic regionIt can produce spines, hairs, lateral shoots or flowers. It is not a single spine or a hole in the epidermis.
RibA longitudinal ridge on the stemIts geometry facilitates volume changes in the hydrated shoot. Rib number may change during growth.
TubercleA distinct projection on the shoot surfaceIt may be associated with an areole. Its arrangement and grooves matter in genus descriptions.
AxilIn mammillaria descriptions, the depression between tuberclesThe flowering region may be separate from the spine-bearing apical portion of the areole.
Radial spineA spine arranged radially near the areole's perimeterIt usually lies closer to the shoot surface. The distinction from a central spine concerns position, not just length.
Central spineA spine developing nearer the centre of the spine clusterIt often projects farther from the surface. It may be straight or hooked.
GlochidA small barbed spine typical of opuntioidsIt detaches easily and penetrates skin. Glochids are not soft hairs that can be touched safely.
Areole hairsTrichomes, often forming a woolly coveringThey should be distinguished from thin or feathery spines. Similar appearance does not demonstrate shared structure.
SegmentA shoot section separated by a distinct constrictionA flat opuntia or schlumbergera segment is part of the stem, not a leaf.
Epidermis and cuticleThe covering tissue and its outer protective layerStomata belong to the epidermis. The cuticle limits water loss but does not seal the plant completely.
Photosynthetic parenchymaLiving cells with chloroplastsIt performs photosynthesis. It usually lies in the outer portion of the shoot cortex.
Water-storage tissuesLiving tissues maintaining a water reserveThey are not an empty cistern. Storage and photosynthetic functions may partly overlap.
Xylem and phloemConducting tissuesXylem carries water and minerals, while phloem mainly carries organic compounds between sites of production and use.
Root collarThe transition zone between shoot and root systemIts position helps assess correct planting depth and the location of damage.
Primary root and lateral rootsThe axis of embryonic origin and its branchesRoots arising from the shoot of a cutting are adventitious. A large root is not necessarily a primary root.
Storage rootA thickened root storing resourcesIt is not an underground stem. Cacti differ in the contribution of roots and shoots to water storage.

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Growth habit and general plant appearance

Table 1 • Growth habit and general plant appearance
Element or termMeaningExamples and explanation
phytonPlant, an element of Greek originAstrophytum combines a reference to a star with the term for a plant. Conophytum is an example outside Cactaceae.
stēlē, columnaColumn or pillarchrysostele, albicolumna. stēlē comes from Greek, columna from Latin.
dendron, arborTreearborescens means becoming tree-like. The element dendro- is Greek.
dumusShrub or scrubdumetorum refers to scrub as a habitat and need not describe the cactus's own habit.
frutexShrubfruticosus, frutescens indicate a shrubby character or its development.
virgultumShrub, scrub, young shootsvirgultus should be read in the context of the original description.
sphaira, globusSpheresphaerocarpus applies globose shape to a fruit, globularis to a spherical shape. Not every element refers to the entire plant.
clava, korynē, rhopalonClubclavatus, claviformis, corynodes. They describe broadening towards the end of an organ.
cylindrus, kylindrosCylindercylindroarticulatus means cylindrically segmented. The element also occurs in Cylindropuntia.
discusDisc, shielddisciformis means disc-shaped. Discocactus illustrates the element's use in a genus name.
caespesTurf, dense clumpcaespitosus means clumping, forming clusters of shoots.
depressusFlattened, loweredDescribes organ proportions. A flattened stem need not be segmented.
prostratus, pendulusProstrate, hangingTwo distinct growth directions. They should not be treated as synonyms.

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Stem, branches and segments

Table 2 • Stem, branches and segments
Element or termMeaningExamples and explanation
caulis, kaulosStemLatin caulis and Greek kaulos denote the shoot axis. fusicaulis refers to a spindle-shaped stem.
truncusTrunkThe correct Latin term for a trunk. The former assignment of “trunk” to stemma was incorrect.
ramusBranchramosus means branched, ramosissimus very strongly branched.
brachiumArmbrachiatus describes arm-like branches.
kladosBranch, offshootheterocladus refers to differences between branches, glaucocladus to their glaucous colour.
fronsLeafy branch or blade, depending on contextlatifrons and acutifrons require checking the organ description. The element alone does not demonstrate typical leaves.
sarmentumLong, slender shootsarmentaceus is associated with this habit. The archive used the inaccurate spelling sarmentium.
articulus, arthronSegment or jointarticulatus means segmented. Arthrocereus contains a Greek element referring to joints between segments.
stemmaWreath, crownThis Greek element remains in the comparison as a correction. It is not a term for a stem or trunk.

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Spines, bristles and hairs

Table 3 • Spines, bristles and hairs
Element or termMeaningExamples and explanation
spina, akanthaSpine, sharp structuretenuispinus, melanacanthus. Etymology does not replace identification of an organ's origin.
aculeusPrickle or sharp outgrowths in older descriptionsaculeatus. In a strict morphological description, a prickle is a surface outgrowth, while a cactus spine has leaf origin.
ankistron, hamus, uncusHookancistrophorus, multihamatus, uncinatus indicate a hooked curvature.
centrumCentrecentrispinus refers to central spines. Position in the areole is distinct from spine length.
acutus, acumenSharp, sharply pointed tipacuminatus indicates a gradually elongated point. The imprecise archival spelling aculius was omitted.
cuspisPoint, sharp endingtricuspidatus means three-pointed. The number refers to a particular feature in the description.
subulaAwlsubulatus describes an awl-shaped form tapering to a sharp end.
acusNeedleacifer, acispinosus refer to a needle-like form.
armaArmamentArmatocereus, albiarmatus. A metaphor for defensive traits.
acinacesShort curved swordacinaciformis, acinacispinus describe a shape resembling a blade.
machairaBlade, knife or swordMachaerocereus and machaeratus illustrate historical use of the element, not a statement of current genus status.
pugioDaggerpugionacanthus, pugionifer refer to the shape or presence of dagger-like structures.
aor, ensis, gladius, xiphosSwordensiformis, gladiispinus, xiphacanthus. These are different roots evoking similar shapes.
hasta, lanceaSpear, lancehastatus and lanceolatus are not interchangeable. In leaf descriptions they denote different outlines.
sarissaLong spearsarrisophorus refers to long, sharp armament.
chaitē, setaBristlechaetacanthus, saetiger, acanthosetus emphasize a thin, bristly form.
pilus, thrixHairLatin pilo- and Greek tricho-. Examples include Pilosocereus and sphaerotrichus.
coma, crinisTuft, hairalbicomus, brevicrinita. A reference to hair does not establish whether the structure described is a trichome or a thin spine.
erion, lanaWooleriophorus, laniceps indicate a woolly appearance of a particular part.
floccus, cincinnusTuft, curlflocculosus, cincinnatus describe covering arrangement, not a separate cactus organ.
pinna, plumaFeatherpinnatus, plumosus. In cacti, a feathery appearance may result from spine structure.
inermisUnarmedClosest in meaning to lacking spines or prickles. subinermis means almost unarmed.
nudusNaked, exposednudiflorus applies nakedness to the flower. It does not automatically mean the entire plant lacks spines.
calvusBald, without coveringcalvescens indicates becoming bald, losing the covering during development.
laevisSmoothlaevigatus refers to a smooth surface. Smoothness and lack of spines are different traits.
glaberHairlessglabrescens means becoming hairless. It concerns hairs, not a general statement of spinelessness.

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Shoot surface, leaves, roots and fruits

Table 4 • Shoot surface, leaves, roots and fruits
Element or termMeaningExamples and explanation
areolaSmall area, in cacti an areolelongiareolatus refers to elongated areoles. An areole is the site where a specialized short shoot develops.
glochisBarb, barbed pointglochidiatus refers to glochids. The singular in a Polish description may be rendered as glochidium.
dermaSkinArgyroderma is an example outside Cactaceae. A surface name does not establish the anatomical covering type.
sperma, semenSeedcardiospermus, dolichospermaticus. The elements may describe shape or other seed traits.
fructus, karposFruitalbifructus, platycarpus apply a trait to the fruit.
baccaBerrybaccifer means producing berries. It is not a synonym for every fruit type.
pulpaPulpduripulpa refers to pulp hardness.
radix, rhizaRootbrunneiradicatus, macrorhizus, polyrhizus. The term may concern root colour, size or number.
rapum, napusTurnipnapiformis describes resemblance to a turnip, usually organ thickening. It is not a general term for a root.
angulus, goniaAnglemultangularis, hexagonus, heptagonus. An angular cross-section may correspond to ribbing, but “angle” and “rib” are not exact synonyms.
costa, pleuraRib, sidepaucicostatus means having few ribs, while pleurocarpus applies ribbing or lateral position to the fruit, depending on the name.
mamma, mastosBreast, projectiongrandimamma, Echinomastus refer to tubercles or resemblance in shape.
papilla, thēlēPapilla, tuberclepapillosus, conothele, Thelocactus. Not every microscopic surface papilla is a large shoot tubercle.
folium, phyllonLeafgrandifolius, rotundifolius, phyllacanthus. The former name Phyllocactus referred to a leaf-like stem, not ordinary leaves.

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The flower and its parts

Table 5 • The flower and its parts
Element or termMeaningExamples and explanation
anthos, flosFlowermicranthus, grandiflorus. The first element is Greek, the second Latin.
kalyx, calyxCalyxGymnocalycium refers to a “naked calyx”. The historical meaning does not replace a modern description of the floral tube and pericarpel.
sepalumSepalcrassisepalus, smaragdisepalus. In many cacti, the boundary between outer and inner tepals is gradual.
petalon, petalumPetaloxypetalus refers to sharply pointed petals. In a description of a continuous perianth, tepals may be the safer term.
stylos, stylusStyle, originally a pillar or stylusbrevistylus, flavistylus describe style length or colour.
graphisStylus, writing toolThe old brachygraphis illustrates a shape metaphor. graphis should not be used as a universal term for the style.
stamen, stēmōnStamenlongistamineus refers to stamens. A stamen includes a filament and anther, so it is not the filament alone.
filamentumThread, including a stamen filamentrubrifilamentosus indicates red filaments. filamenta is plural.
anthera, stigma, ovariumAnther, stigma, ovaryThree supplementary anatomical terms. The anther produces pollen, the stigma receives it, and the ovary contains ovules.

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Former names in the supplied books

Examples of updates using POWO, checked 9.10.2026
Former nameAccepted nameRelationship and significance
Zygocactus truncatusSchlumbergera truncataChanged generic combination. We retain the former name when discussing the book. [2-WWW]
Chamaecereus silvestriiEchinopsis chamaecereusThe accepted name follows current POWO. The older combination helps find the earlier description. [2-WWW]
Mammillaria wildiiMammillaria crinita subsp. wildiiIt has subspecies rank in the adopted treatment. [2-WWW]
Hylocereus undatusSelenicereus undatusTransfer to another genus does not mean a different species cultivated as a rootstock. [2-WWW]
Dolichothele longimammaMammillaria longimammaA former genus found in literature describing elongated tubercles. [2-WWW]
Neomammillaria gaumeriMammillaria heyderi subsp. gaumeriThe name from Britton and Rose's monograph now corresponds to a subspecies in POWO's treatment. [2-WWW]

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Exhibitions and the scale of development

PTMK membership in historical tabulations
Date of tabulationMembersData scope
18 November 1966388Total excluding Bydgoszcz data. Includes 25 people from provinces where branches had not been constituted.
Second General Congress, June 1969119813 branches. The authors separately record 15 corresponding members from other countries.
Third General Congress, 1972About 30010 branches. The source gives an approximate value.

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Water loss, sugars and mannitol

Osmotic pressure of photosynthetic parenchyma after two temperature regimes (MPa)
Material identified in the study30/20°C, day/night10/0°C, day/night
Opuntia ficus-indica0.70 ± 0.020.80 ± 0.01
Opuntia humifusa0.88 ± 0.051.23 ± 0.06
Opuntia streptacantha0.71 ± 0.020.83 ± 0.03

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Seasonal resistance on the Canadian prairie

Seasonal change in segment tolerance from Saskatchewan
Date and materialWater (% fresh mass)LT₀, no visible damageLT₅₀, tissue damage
6 IX 1985, mature segments O. fragilis84.3%−7°C−10°C
10 X 1985, O. fragilis71.9%−50°Cbelow −70°C
10 XII 1985, O. fragilis69.6%−50°Cbelow −70°C
8 IV 1986, O. fragilis72.5%−30°Cbelow −60°C
18 VI 1986, young segments O. fragilis91.0%not determineddeath after freezing at −3.5°C
10 XII 1985, O. polyacantha67.6%−40°C−55°C

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Twenty populations of Opuntia fragilis

LT₅₀ after acclimatization and improvement relative to 25/15°C
Material provenanceLT₅₀ (°C)Tolerance improvement (°C)
Peace River, Alberta−39.020.0
Fort St. John, British Columbia−44.324.3
Goose Lake, Saskatchewan−41.123.4
Cache Creek, British Columbia−34.914.9
Bird River, Manitoba−22.617.2
Vernon, British Columbia−30.111.1
Hornby Island, British Columbia−19.112.1
Lethbridge, Alberta−28.319.0
Virden, Manitoba−48.919.9
Minot, North Dakota−40.217.2
Plymouth, Washington−20.611.1
Eau Claire, Wisconsin−25.518.8
Kaladar, Ontario−20.415.4
Thermopolis, Wyoming−28.617.6
Salt Lake City, Utah−25.017.0
Leadville, Colorado−22.416.4
Princeton, Colorado−15.614.2
Russell, Kansas−23.319.3
Loa, Utah−23.613.6
Albuquerque, New Mexico−27.619.6

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Shoots, roots, fruits and meristems respond differently

LT₅₀ of organs of Opuntia ficus-indica clone 1279 (°C)
Organ and ageAfter 20/10°CAfter 30/20°C
One-year segment−7.6 ± 0.2−6.7 ± 0.2
Six-week segment−7.8 ± 0.2−6.5 ± 0.1
Six-week fruit−6.3 ± 0.3−6.3 ± 0.2
Six-week root−6.5 ± 0.2−6.4 ± 0.2

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Shoots, roots, fruits and meristems respond differently

Tissue LT₅₀ after 100 days at 10/5°C (°C)
Material in the publicationPhotosynthetic parenchymaAreole meristem
Nopalea cochenillifera−9.1 ± 0.4−11.3 ± 0.4
Opuntia robusta−8.8 ± 0.4−11.2 ± 0.4
Selenicereus megalanthus−6.5 ± 0.3−7.4 ± 0.4

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Field studies and a real winter

Damage to three-year-old opuntia segments after frost in Mendoza (%)
Material named in the publicationPlainDuneMean
O. paraguayensis0.00.70.3
O. robusta3.00.92.0
O. spinulifera3.76.55.1
O. ficus-indica “San Rafael”19.210.614.9
O. ficus-indica “Cuenca”25.723.624.6
O. ficus-indica “San Juan”52.616.134.4

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Differential diagnosis table

Physiological symptom and checking its cause
ObservationWhat to checkCourse of action
Pale patch after a change of positionRelationship to exposure, side of the stem, temperatureGradual acclimatisation and observation of new growth
Narrow, elongated growthTypical growth form and lighting historyImproved light suited to the growth stage
Wilting in a wet potRoot condition, drainage, duration of moisture retentionRestoring suitable root conditions and distinguishing infection
Chlorosis after intensive fertilisationEC, pH and root viabilityIdentifying the problem before further fertilisation
Watery tissue after coolingTemperature history and expansion of damageIsolating the plant and assessing lasting damage and secondary rot

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Products checked in official documents

Document status on 9.10.2026. Use depends on the full label
Product and userWhat the document statesRelevance to cactus cultivation
Sanium AL
Non-professional
Flupyradifurone, a ready-to-use spray product. Ornamental plants indoors and under cover, against pests including aphids, scale insects and greenhouse whitefly. [10-WWW]Do not extend this scope to root drenching. Outdoor uses differ. The full label applies.
Mospilan 20 SP
Professional
Acetamiprid. The label dated 11.08.2025 includes specified minor uses in ornamental plants under cover and in the ground. [10-WWW]A product for professional users. The spraying scope does not include arbitrary drenching of the root ball.
Magnicur Energy
Non-professional
Propamocarb and fosetyl. The label dated 24.02.2026 includes Phytophthora disease and diseases caused by Pythium in ornamentals, under specified conditions and at specified stages. [10-WWW]It is not a universal product for every rot. The cultivation site, plant stage and diagnosis must be checked for the particular method.
Actellic 500 EC
Professional
Pirimiphos-methyl. The current label concerns disinsection of warehouses and protection of specified cereal grain. [10-WWW]The historical recommendation to use it against cactus pests does not match the current scope of this label.
Kaptan Gold 80 WG
R-101/2021
The register gives 30.09.2026 as the deadline for using existing stocks. [10-WWW]The deadline had passed by the verification date. This does not mean that all other captan-containing products have been withdrawn.

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Historical names still found in the current register

Products still registered. Current scope and restrictions
Product and userCurrent useComment on the historical book
Basamid
Professional. Fumigation training required
Dazomet. Soil or substrate disinfection before planting and sowing, within the scope specified by the label. [10-WWW]The product remains in the register. This is a specialist substrate preparation treatment, rather than treatment of plants in a household pot.
Siarkol 80 WP
Professional
Sulphur, a fungicide. The label includes powdery mildew of ornamental plants. [10-WWW]The current formulation and its use are given. Historical sulphur use against “red spider” has not been carried over to the scope of this label.
Kaptan Zawiesinowy 50 WP
Professional
Captan. The label dated 7.11.2025 includes apple and pear scab and, in permanent greenhouses, scab and leaf spots of ornamental trees and shrubs. For annual ornamental plants, it includes grey mould. [10-WWW]Do not equate it with Kaptan Gold 80 WG or an old general recipe for cacti. The range of crops and diseases listed on this label applies.
Miedzian 50 WP
R-134/2015
Professional and non-professional uses
Copper oxychloride. Uses include apple scab and fire blight. The documents specify the scope and method of use separately for professionals and amateurs. [10-WWW]The brand remains in the register. Orchard and vegetable uses are not automatically extended to control of cactus rots.

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Historical products not adopted in the recommendations

Historical names whose old recipes must be rejected
Names in historical guidesComparison resultConclusion
Benlate, Fundazol, Dithane M-45, Novozir MN 80No current products under these exact names were found in the register checked. [10-WWW]Do not reproduce historical concentrations or mixtures. Any new product is assessed according to its own authorisation.
Ekatin, Anthio, Metasystox R, Rogor, Ultracid 40 ECThese exact names are absent from the list checked. [10-WWW]Historical mealybug or mite control is not a current application instruction.
DDT and mercury products described historicallyThey have not been adopted in the recommendations. Fleischer and Schütz already noted restrictions on older products in their 1978 edition. [Fleischer and Schütz 1978, pp. 188–193]The history of cultivation advice is retained without reproducing obsolete practice.
General names “sulphur”, “copper”, “captan”The substance name alone does not specify a product or its current scope. [10-WWW]Do not label an entire substance as authorised or withdrawn on the basis of one brand.

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How to compare the eight diagrams

Tubercle traits and botanical examples
Trait comparedExampleBasis of interpretation
Short-conicalMammillaria mammillarisMorphological description [Britton and Rose 1923, p. 70], current name [2-WWW]
ElongatedMammillaria longimammaMorphological description [Britton and Rose 1923, pp. 61–63], current name [2-WWW]
Cylindrical and roundedMammillaria bocasanaMorphological description [Britton and Rose 1923, p. 147], current name [2-WWW]
Slender and strongly taperingMammillaria longimamma, comparison of elongationMorphological description [Britton and Rose 1923, pp. 61–63], current name [2-WWW]
Blunt-tippedMammillaria heyderi subsp. gaumeriMorphological description [Britton and Rose 1923, pp. 72–73], current name [2-WWW]
Angular, approaching pyramidalMammillaria polyedraMorphological description [Britton and Rose 1923, pp. 102–103], current name [2-WWW]
Laterally flattenedMammillaria compressaMorphological description [Britton and Rose 1923, pp. 90–92], current name [2-WWW]
Densely arrangedMammillaria evermannianaMorphological description [Britton and Rose 1923, p. 97], current name [2-WWW]

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Two different angles

Angle of the stem axis to the horizontal in the 1980 study [Ehleringer et al. 1980, Table 1. SD denotes standard deviation]
Taxon and locationNumber of plantsMean ± SD
C. cinerea, Paposo5044.6° ± 6.3°
C. cinerea, Taltal5065.6° ± 8.6°
C. cinerea subsp. columna-alba, Pan de Azúcar10051.9° ± 8.6°
C. gigantea, Paposo5042.1° ± 8.8°

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Category counts

Categories according to the Results section of the 2024 publication [Villalobo-Lopez et al. 2024, pp. 6–7]
CategoryNumber of taxaShare of 37 assessments
CR · Critically Endangered1437.8%
EN · Endangered1437.8%
VU · Vulnerable616.2%
LC · Least Concern38.1%
Total CR + EN + VU3491.9%

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What differs between close relatives

Comparison of growth forms and flowers [de Vos et al. 2025][Korotkova et al. 2010]
TraitCalymmanthiumLymanbensonia
HabitBranched shrubs or treesSmaller plants, erect, spreading or hanging
StemSeveral thin, wing-like ribsFlattened or angular, depending on species
SubstrateTerrestrial growthTerrestrial, rock-dwelling or epiphytic growth
Flower budHidden inside the short-shoot covering, which opens before floweringMore typical architecture, with an exposed developing perianth

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Two historical names in one current species

Selected chromosome count results in the 2019 paper [Las Peñas et al. 2019] [2-WWW]
Current nameIdentification of material in the paper2nNumber of x sets
T. alexanderiT. alexanderi222
T. bonnieaeT. bonnieae444
T. articulatusvar. strobiliformis777
T. articulatusvar. articulatus and var. oligacanthus888
T. aoracanthusT. aoracanthus24222
T. aoracanthusT. paediophilus31929

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Sixty labelled stem segments

Successive runoff episodes. Distances as mean ± standard deviation [Almirón et al. 2019]
EpisodeRainfall and durationMean displacement
First35 mm in 2 days480.4 ± 210.0 m
Second100 mm in 4 days103.0 ± 138.9 m
Third20 mm in several hours33.2 ± 52.3 m

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Two circular sequences

Assembled sequences of two isolates [Oses Pedraza et al. 2023] [Bartholomäus et al. 2024]
IsolatePublication yearGenome lengthGC content
Arthrobacter sp. ATA00220233 904 590 base pairs65.9%
Cellulomonas sp. ATA00320244 084 881 base pairs73.8%

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The leaf's full-day balance

Daily components of leaf CO₂ uptake. Conditions 25/15°C, day/night of 12 h each [Nobel and Hartsock 1987]
SpeciesDrought [days]Day [mmol m⁻²]Night [mmol m⁻²]
Pereskiopsis porteri01087
Pereskiopsis porteri7672
Pereskiopsis porteri14135
Quiabentia verticillata (as Q. chacoensis)01299
Quiabentia verticillata72416
Quiabentia verticillata1402
Austrocylindropuntia subulata01112
Austrocylindropuntia subulata7624
Austrocylindropuntia subulata14015

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Two species, two responses

Germination after 30 days. Five replicates of 25 seeds per treatment [Jara-Peña et al. 2024]
Species and origin12 h of light per dayContinuous darkness
Austrocylindropuntia floccosa, Apurímac28.0%32.8%
Cumulopuntia ignescens, Arequipa36.0%0%

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Three levels of the result

Measures of ant–cactus association [Braun and Lortie 2025][Lynch and Savage 2023]
LevelWhat was measuredMeaning
VisitsNumber and identity of antsResource access and activity on the plant
DefenceBug numbers or response to test preyHerbivore pressure or defensive behaviour
ReproductionNumber of developed seeds per fruitOne component of reproductive outcome

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Force and strength describe different traits

Stem-segment junction tension. Medians in the experiment by Mylo and colleagues (2022) [Mylo et al. 2022]
Material studiedMaximum force [N]Tensile strength [MPa]
Cylindropuntia bigelovii14.41.41
Opuntia ficus-indica, younger lateral segment51.23.84
Opuntia ficus-indica, older lateral segment1532.70

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Wound periderm

Course of healing under the experimental conditions of 30/20°C [Mylo et al. 2020]
Observation stageTissue change
First daysEdge curling and a lipophilic barrier at the wound surface
Around the first weekVisible pale layer of dead cells and development of wall changes
Subsequent weeks, through day 31Organisation of wound periderm and remodelling of tissues beneath the surface

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Directions in wood

Levels of wood organisation in Cylindropuntia imbricata [Morankar et al. 2024]
Structural scaleVisible elementsScope of information
Stem skeletonOpenings and connected wood bandsGeometry of the whole structure
Tissues and cellsVessels, fibres, raysDirection and type of wood organisation
Cell wallLayers and pitsLocal structure and response to pressure

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How many plants the camera detects

Correctly detected S. wrightiae relative to 480 plants in the ground census. Data from Bates et al. (2022), Table 2. Percentages were calculated from total counts. [Bates et al. 2022, Table 2, p. 7]
Flight heightCorrectly detectedProportion of ground censusMissed
10 m18338,1%297
15 m8918,5%391
20 m469,6%434

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Inhibition of two enzymes

IC50 concentrations in enzyme tests, in µg/mL. Mean ± SD, three replicates described by the authors as biologically independent. A dash indicates the absence of a value in the source table. [Heraiz et al. 2026, Table 2, p. 8]
Material testedAChEBACE-1
Spines, methanolic extract0,362 ± 0,0120,425 ± 0,014
Stem, methanolic extract0,603 ± 0,0190,939 ± 0,031
A. oryzae culture, ethyl acetate extract0,255 ± 0,0080,550 ± 0,018
A. oryzae culture, subsequent methanolic extract3,180 ± 0,1041,882 ± 0,062
Donepezil, reference substance for AChE0,167 ± 0,006—
Curcumin, reference substance for BACE-1—0,350 ± 0,012

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Cooler air beside the stem surface

Daytime microclimate beside Eulychnia breviflora. Means from four sensor pairs, Table 1 of Stanton and colleagues. [Stanton et al. 2014]
ParameterWith epiphytesWithout epiphytes
Mean daytime temperature15,1°C17,3°C
Mean of daily maximum temperatures20,9°C25,1°C
Mean daytime VPD0,69 kPa1,02 kPa
Mean of daily maximum VPD values1,54 kPa2,33 kPa

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Seeds after passage through the digestive tract

Germination of surviving Eulychnia acida seeds in the experiment by Cares and colleagues. 100 seeds per group, about 26°C, photoperiod 14:10 h. [Cares et al. 2018]
Seed originGerminationMeaning of the comparison
Directly from fruits52%Control group
Recovered after goats60%No demonstrated difference from the control
Recovered after guanacos13%Lower than in the control

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The same seed batch under four conditions

Germination of seeds from the Santa Cruz population after four months. Mean ± standard error, three replicates of one hundred seeds each. [Jaramillo et al. 2010, methods on p. 22 and Table 2 on p. 24]
PreparationSubstrate in the dishGermination [%]
Soaking for 24 h at 25°CMoist filter paper52,0 ± 10,2
Soaking for 24 h at 60°CMoist filter paper47,3 ± 8,1
No soakingMoist soil7,3 ± 3,0
No soakingMoist filter paper66,7 ± 8,6

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Crystals and fibres in a precisely defined location

Traits of the lineages compared in the 2019 studies. [Vázquez-Sánchez et al. 2019, Vázquez-Sánchez et al., pp. 409, 412, 416–417. De la Rosa-Tilapa et al., online version, pp. 2–4] [De la Rosa-Tilapa et al. 2019, Vázquez-Sánchez et al., pp. 409, 412, 416–417. De la Rosa-Tilapa et al., online version, pp. 2–4]
GenusSeed microsculptureHypodermal crystalsFibres in cortical bundles
TurbinicarpusStriatePrismaticAbsent in the material studied
RapicactusPapillateConcentric drusesAbsent in the material studied
KadenicarpusSmoothPrismaticPresent in the material studied

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Two lineages in a mountain landscape

Site elevations in the data of Donati and colleagues. Ranges concern the dataset from the 2017 paper. [Donati et al. 2017, online version, pp. 5–7]
Group in the studySite elevationBand with the most taxa
Rapicactus1360–3120 m above sea level1400–1800 m above sea level
Turbinicarpus as treated by the authors307–2185 m above sea level1000–1400 m above sea level

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A papery ribbon and a corky structure

Fibre-wall thickness in selected radial spines, in µm. Mean ± standard deviation. [Mosco 2009, Table 2, pp. 122–123]
Taxon studiedSpine descriptionFibres in the deeper portionThin-walled fibres
T. valdezianusWoody structure, feathery epidermis4,1 ± 0,9Not found
T. schmiedickeanus subsp. flaviflorus (T. flaviflorus in the paper)PaperyNot found1,5 ± 0,3
T. schmiedickeanus subsp. rioverdensis (T. rioverdensis in the paper)Corky, with fissures3,1 ± 0,91,0 ± 0,2

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Multiplication and symptom frequency

Shoot number and explants with hyperhydricity symptoms in T. valdezianus. Comparison from Table 1 of the publication, described as an assessment after 12 weeks. [García Osuna et al. 2011, Table 1, p. 178]
Additive to the kinetin mediumShoots per explant, meanExplants with symptoms
Control without the tested additives3,55 b25%
Paclobutrazol, 3,4 × 10−4 M18,00 a0%
Prohexadione-calcium, 10−4 M3,25 b50%
Salicylic acid, 10−4 M1,15 b75%
Benzoic acid, 10−4 M1,20 b75%

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Different shrubs, different conditions

Abundance over a total area of 490 m². Data from Table 2 of the 2026 study. [Jiménez-Sierra et al. 2026, Table 2, p. 5]
MicrohabitatAreaProportion of areaIndividuals
Beneath vegetation281,57 m²57,5%296
Open space208,43 m²42,5%84
Total490 m²100%380

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Buds remaining underground

Diagnostic features of the underground stem and root of Lophophora williamsii. Histological study of five plants. [Terry and Mauseth 2006, pp. 582–589, Table 2]
Tissue or structureUnderground stemRootDevelopmental significance
Outer parenchymaCortex with xylem and phloem bundlesIn older portions, a zone of secondary phloem parenchyma without a cortical bundle networkThe similar appearance of the swollen structure has different anatomical foundations
Organ centreParenchymatous pith without xylem elementsExpanded metaxylem and inner secondary xylem with scattered conducting elementsAllows organ identity to be determined in section
Areoles and their meristemsLiving remnants can persist beneath the soil surfaceNo areolesRetained stem buds produce new branches

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Clusters among empty plots

Random transects in six known populations of Lophophora williamsii. Each transect occupied 100 m². Data from May and July 2019, from Table 6 of the publication. [Ermakova et al. 2021, p. 156, Table 6]
Region and siteTransects with plants / all transectsNumber of plantsDensity on transects (ha⁻¹)
South, site 14 / 2771262,96
South, site 23 / 3173235,48
South, site 31 / 2653203,85
West, site 41 / 1425178,57
West, site 51 / 1826144,44
West, site 64 / 546920,00

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Changing proportions with water availability

Seedling growth after six months of the experiment. Both light levels combined. Mean ± standard deviation. [Martínez-Berdeja and Valverde 2008, Table 4, p. 1770]
Substrate hydration relative to the reference valueRelative growth rate, g g⁻¹ day⁻¹Root-to-shoot dry-mass ratioRatio of relative root and shoot growth rates, K
25%0,007 ± 0,0021,106 ± 0,3532,775 ± 0,895
60%0,009 ± 0,0030,774 ± 0,3881,51 ± 0,256

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Medium additives and the quality of new shoots

Response of Obregonia to additives in the common CM medium. Mean ± standard deviation. Data pooled for two explant types. [Cardarelli et al. 2010, Table 3, p. 34]
Medium treatmentShoots per explantExplants with callus, %Spontaneous rooting, %
CM1,0 ± 0,5——
CM + putrescine, 1 mM2,4 ± 0,616,7 ± 1,133,3 ± 3,7
CM + AgNO₃, 10,2 µM1,2 ± 0,315,0 ± 1,014,6 ± 1,3
CM + CPPU, 12,1 µM5,0 ± 0,436,8 ± 4,2—

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Numbers of flowers, fruits and seedlings

Flowering and recruitment in one population in Querétaro. The initial census included 468 plants in an area of approximately 20 × 30 m. Searches for new seedlings were conducted twice a year. [Briseño-Sánchez et al. 2022, Table 1]
YearNumber of flowersFlowers forming fruitsNew seedlings
201546244,3%21
201656222,2%21
201778326,3%31
201883544,5%30
201911562,6%5
202016225,9%1
202142332,8%18

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Seeds stored for four years

Comparison of seed batches in the experiment conducted in 2018. Each batch comprised 50 seeds in ten dishes. Germination means the observed emergence of the radicle, and survival concerns established seedlings at the end of the short laboratory observation. The last column gives the mean ± standard error. [Briseño-Sánchez et al. 2024, pp. 5–7, Fig. 3–4]
Collection year and storageSeeds in the sampleObserved germinationSeedling survival
2018, from the year of the experiment5062%61 ± 5,92%
2014, approximately four years5052%58 ± 4,81%

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A seed from a rocky canyon

Comparative characters of the taxa described. Botanical ranges from Arias and Sánchez-Martínez (2010), table 1. [Arias and Sánchez-Martínez 2010, p. 621, Table 1]
TaxonStem, height × diameterSpinesPerianth colour
S. disciformis subsp. disciformis3–12 × 3–9 cm1–5, flexible, shed earlyWhite to yellowish white
S. disciformis subsp. esperanzae2,5–4,5 × 4–5 cm1–5, flexible, shed earlyIntense magenta
S. corregidorae18–23 × 8–12 cm3–5, longer and persistentYellow

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Air in the aril and coat cells

Return to the surface after controlled submersion to a depth of 8 cm. 40 different seeds per taxon, mean ± standard deviation. [Camacho-Velázquez et al. 2018, pp. 91, 94]
TaxonArilTime to return to the surface
S. disciformis subsp. disciformisPresent2,61 ± 0,53 s
S. disciformis subsp. esperanzaePresent1,74 ± 0,65 s
S. corregidoraeAbsent4,72 ± 1,55 s

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Germination at a specified temperature and light regime

Germination at 25°C with 12 hours of light, on moist filter paper. Eight replicates of 30 seeds per taxon, observed for 30 days. Mean ± standard deviation. [Camacho-Velázquez et al. 2018, pp. 91, 94]
TaxonArilGerminationMean germination time
S. disciformis subsp. disciformisPresent97,08 ± 3,75%5,29 ± 0,04 days
S. disciformis subsp. esperanzaePresent92,08 ± 7,75%5,27 ± 0,06 days
S. corregidoraeAbsent96,67 ± 4,36%5,31 ± 0,06 days

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Measuring nectar at a specified time

Tissue structure and refractometric nectar reading. Nectar collected at 15 hours in February 2018 from 15 flowers of different individuals of each taxon in the UNAM collection. The publication does not unambiguously define the measure of variation following the ± sign. [Camacho-Velázquez et al. 2019, pp. 102–104]
TaxonEpidermal cellsSecretory parenchyma layersNectar reading
S. disciformis subsp. disciformisSlightly convexApproximately 10–1222,09 ± 7,42°Brix
S. disciformis subsp. esperanzaeRelatively flatApproximately 8–98,65 ± 3,98°Brix
S. corregidoraePapillate convexApproximately 7–816,12 ± 4,48°Brix

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Which processes had the greatest effect

Process elasticities in the P. bradyi model. Values from table 2 of the 2026 paper.
ProcessElasticity of λ
Survival0,39
Stem growth in the visible state0,24
Root elongation and exit from burial0,17
Growth after the buried state0,15
Entry into the buried state−0,01

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Twenty years in one Utah plot

Scopes of three studies. Demographic data and habitat comparisons represent separate parts of the analyses.
SpeciesMaterial and periodMain observations
P. bradyi965 plants, three plots, a 31-year databaseRelationship of survival to diameter and transition to the buried state
P. nigrispinusFour demographic plots, censuses in 2016–2019 and 2022Loss of recorded plants, no new seedlings detected
P. winkleri107 plants in one plot, 1995–2014Effects of trampling and weather, reproduction concentrated in some specimens

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Cytokinin, apical dominance and successive cycles

Selected multiplication results from 2002. Number of shoots per initial explant, mean ± standard error. [Pérez-Molphe-Balch and Dávila-Figueroa 2002, Table 1–2, pp. 74–75]
TreatmentShoots after 60 daysShoots after 180 days
P. aselliformis, apex
BA 4,4 µM, sucrose 30 g/L
13,23 ± 1,41122,27 ± 9,05
P. aselliformis, apex
BA 8,8 µM, sucrose 30 g/L
13,70 ± 1,37128,07 ± 9,16
P. strobiliformis, disc
BA 4,4 µM, sucrose 50 g/L
12,37 ± 1,21119,33 ± 9,66
P. strobiliformis, apex
BA 4,4 µM, sucrose 30 g/L
6,43 ± 0,40136,33 ± 12,6

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The apex and an areole-bearing segment give different results

Micrografting of P. aselliformis onto O. ficus-indica. Mean ± standard error in the 2016 experiment. [Badalamenti et al. 2016, pp. 3–4, Table 1]
MeasureApical scionSubapical segment
Scion length5 mm3 mm
Number of micrografts30270
Successful micrografts97 ± 0,7%81 ± 1,2%
New shoots per scion1,0 ± 02,7 ± 0,3
Length after 3 weeks31 ± 0,6 mm28 ± 0,4 mm
Length after 6 weeks38 ± 0,3 mm35 ± 0,5 mm

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Five or six flowering waves

Flowering of Neolloydia conoidea in Tolimán. Weekly observations. Mean fruit set ± standard error, the proportion of flowers forming fruits. Data from Arroyo-Pérez et al. 2019, table 1 and sample description. [Arroyo-Pérez et al. 2019, pp. 581–583, Table 1]
YearPlants in the analysisFlowering wavesRecorded flowersFruit set
201514669300,67 ± 0,022
201614259070,58 ± 0,023

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Pollen from other plants and the number of seeds formed

Controlled pollination of Neolloydia conoidea in Tolimán. 20 flowers per treatment. Fruit set is the percentage of flowers forming fruits. Seed set is the mean proportion of ovules forming seeds according to table 3 of Arroyo-Pérez et al. 2021. [Arroyo-Pérez et al. 2021, pp. 32, 35, Table 3]
TreatmentFruit setSeed set
Insect access, control90%0,73
Hand cross-pollination95%0,96
Hand self-pollination0%0
Isolation without pollination0%0

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Multiplication, shoot numbers and length

Multiplication of Ortegocactus macdougallii after 110 days, with NAA 4,4 µM in every treatment. Means ± standard error. Eight vessels containing six explants each per treatment, with the experiment repeated twice. Data from Arellano-Perusquía et al. 2013, table 2. [Arellano-Perusquía et al. 2013, pp. 161, 163, Table 2]
BA (µM)Shoots per explantShoot length (mm)Fresh mass (mg)
13,35,67 ± 0,494,78 ± 0,17340,48 ± 47,06
19,95,83 ± 0,653,93 ± 0,26388,62 ± 36,79
26,65,33 ± 0,723,67 ± 0,15327,64 ± 26,17

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A separate stage of root formation

Rooting of Ortegocactus macdougallii shoots after 50 days. Means ± standard error. Eight vessels containing three explants each per treatment. IBA concentrations are given in µM in accordance with the methods and results descriptions. Data from Arellano-Perusquía et al. 2013, table 4. [Arellano-Perusquía et al. 2013, pp. 161, 164, Table 4]
MediumForming roots (%)Number of rootsRoot length (mm)
¼ MS, without regulators58,34 ± 8,344,17 ± 0,4811,32 ± 1,95
½ MS, without regulators37,49 ± 11,685,00 ± 1,379,96 ± 3,05
½ MS + IBA 1,6 µM45,83 ± 6,116,33 ± 0,778,99 ± 0,91
½ MS + IBA 4,9 µM75,03 ± 5,459,92 ± 1,918,37 ± 1,56

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What the habitat model represents

Potential habitat area in three models. Values from table 2 of Solórzano et al. (2026).
Variable setPotential areaModel assessment in the publication
Climate34 469,25 km²Lowest AICc and highest TSS of the three models.
Soil77 082,23 km²Broadest projection, including beyond the confirmed range.
Climate and soil37 519,32 km²Model preferred by the authors for its agreement with field observations and highest pROC.

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Accepted names and the 2026 expansion

Oehmea. Atlas classification and combinations from 2026. POWO status checked on 11 October 2026.
Name accepted in the atlasName in older accountsContext
Oehmea beneckeiMammillaria beneckeiAccepted by POWO. Buxbaum's 1951 combination.
Oehmea sphacelataMammillaria sphacelataAccepted by POWO. Guiggi's 2022 combination.
Oehmea tonalensisMammillaria tonalensisAccepted by POWO. Guiggi's 2022 combination.
Mammillaria napinaMammillaria napina2026 proposal. Oehmea napina.
Mammillaria zephyranthoidesMammillaria zephyranthoides2026 proposal. Oehmea zephyranthoides.

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Meristem size and stem size

Meristem dome width in eight stages of the San Ángel material studied. Mean ± standard deviation, from table 2 of López-Ruiz et al. (2026). [López-Ruiz et al. 2026, p. 1333, Table 2; stage characteristics, pp. 1328, 1330–1334]
Age after germinationDome widthStage characteristic
1 day102,0 ± 30,2 µmClosed cotyledons, two recognisable layers.
3 days94,2 ± 23,7 µmTubercle primordia, three meristem layers.
7 days140,2 ± 17,0 µmFirst visible spines.
15 days128,1 ± 23,9 µmAt least two tubercles with areoles.
30 days137,0 ± 24,9 µmDifferentiation of the areole-covered part of the stem.
90 days212,1 ± 34,0 µmMeristem enlargement and spine stiffening.
1 year224,7 ± 22,4 µmDistinct spine pattern.
4 years771,7 ± 64,5 µmFlowering greenhouse plants.

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Different Mammillaria occupy different places

Spatial associations in the study by Peters et al. (2008). “Positive” means more plants than expected in a given microhabitat, and “negative” means fewer. Accepted names follow POWO. Historical supertexta groups are retained separately. [Peters et al. 2008, p. 596, Table 1] [2-WWW] [2-WWW]
Plant studiedShrubsStones ≥20 cm
M. dixanthocentronAssociation not significant in the testPositive
M. hernandeziiNegativeStones of this size were absent at the sites
M. kraehenbuehliiPositivePositive
M. supertexta (called M. lanata in the paper)NegativeNegative
M. napinaNegativeAssociation not significant in the test
M. pectiniferaAssociation not significant in the testPositive
M. solisioidesNegativeNegative
M. supertexta (under the same name in the paper)NegativePositive

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From a released seed to a seedling after the dry season

Serotiny in Mammillaria pectinifera. Estimates for 238 m² in Texcala
Quantity1997–19981998–1999
Seeds released from retained fruits35 50832 259
Seeds released from ejected fruits07634
All released seeds, estimate35 50839 893
Seedlings surviving the following dry season, estimate1795
Seedling establishment success, ‰0,472,38

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Medium, light and the importance of a specific stage

Snowcap in vitro. Medium stages according to table 1 and the results of the 2023 paper.
StageRegulators in MSSucroseEffect in the experiment
Shoot maintenance, MSM-1NAA 0,5 mg/l + KIN 0,3 mg/l20 g/lGrowth and material maintenance
Offset stimulation, MSM-2NAA 0,5 mg/l + KIN 0,3 mg/l + GA₃ 5 mg/l for 3 days20 g/lSubsequent growth without GA₃
Rooting, MSM-3NAA 0,5 mg/l + KIN 0,3 mg/l + ancymidol 5 mg/l10 g/lRoot development on offsets

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Flowers require different preparation

Snowcap. Flower initiation in vitro after plant preparation and paclobutrazol treatment
Paclobutrazol, mg/lExplants initiating flowers, %Flowers per explant
00 ± 00 ± 0
0,30 ± 00 ± 0
0,50 ± 00 ± 0
166 ± 61,40 ± 0,16
321 ± 50,37 ± 0,10
511 ± 40,24 ± 0,09

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Soil and landforms

Selected site associations with soil and landscape map categories. Results of the frequency analysis in Aquino and co-authors (2021). [Aquino et al. 2021, pp. 4524–4525, Fig. 3]
SpeciesCategories with an excess of sites
E. bokeiCalcaric regosols
E. pachyrhizaCalcaric xerosols and foothill zone
E. pulchraLuvic kastanozems
E. spinosiorLuvic kastanozems and fold mountains
E. polycephalaFold mountains

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Prism, druse, spherulite

Selected biomineral forms and examples directly shown. Plant names follow the current atlas treatment. [De la Rosa-Tilapa et al. 2022, pp. 3–7, Fig. 2, 4–6]
FormTissue and exampleObserved characteristic
PrismsCortex of Epithelantha micromerisCrystals with distinct flat faces
DrusesCortex of Kroenleinia grusonii (called Echinocactus grusonii in the paper)Clusters with projecting tips
Crystal sandCortex of Mammilloydia candida (called Mammillaria candida in the paper)Fine particles up to approximately 5 µm
SpherulitesPith of Mammilloydia candidaRadial arrangement and concentric bands
Silica bodiesHypodermis of some Echinocactus and Ferocactus studiedAmorphous deposits without birefringence

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Unusual polymers in former Escobaria

Chemical identification of selected seed samples. The scope of the results corresponds to the material identifications in Chen and co-authors (2013). [Chen et al. 2013, pp. 203–208, Fig. 4–5]
Current nameName used in the studyResult
Pelecyphora dasyacanthaEscobaria dasyacanthaPredominance of 5H. Almost homogeneous type in the NMR spectrum
Pelecyphora lloydiiEscobaria lloydii5H and S units in chemical analysis products
Pelecyphora zilzianaEscobaria zilziana5H and S, in proportions of approximately 63:37 in the analysed spectrum
Pelecyphora sneedii subsp. sneediiEscobaria albicolumnariaTypical G/S lignin
Pelecyphora viviparaEscobaria vivipara, arizonica and neomexicana materialTypical G/S lignin
Epithelantha micromeris and E. greggiiThe same species namesLignin undetectable in the tests used

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Cell size and wall thickness

Selected mean spine cell-wall thicknesses in the study by Řepka and Gebauer (2012). Measurements were made on hydrated preparations. Unit µm. [Řepka and Gebauer 2012, p. 141, Table 2]
Material studiedEpidermisCore fibres
P. abdita (E. abdita)3,02,4
P. missouriensis, Montana sample (E. missouriensis subsp. missouriensis)4,53,7
P. missouriensis, Arizona sample (E. missouriensis subsp. navajoensis)3,64,0
P. minima (E. minima)3,63,3
P. hesteri (E. hesteri)4,23,8

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Seeds from different zones were transferred to common conditions

Germination under the experimental conditions. Age means the estimated age of the tubercle zone, and the percentage is the mean reported in the publication. [Jiménez-Sierra et al. 2022, pp. 364–365, Tables 1–2]
MaterialEstimated zone ageSeeds / dishesMean germination
Fresh fruitscurrent year's cohort100 / 1013%
Zone 12,7 years100 / 1056%
Zone 27,3 years100 / 1051%
Zone 313,3 years80 / 852%
Zone 419,8 years19 / 373%
Zone 523,9 years39 / 340%

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Self-pollen was often arrested in the style

Response to hand self-pollination. Microscopy and fruit set were assessed in separate flowers. n denotes the size of each sample. [Martínez-Peralta et al. 2014b, p. 535, Table 1]
Species under the current treatmentOvaries with pollen tubes (%) / nFlowers setting fruit (%) / n
A. agavioides25 / 286,9 / 29
A. bravoanus33,3 / 30 / 5
A. fissuratus7,7 / 391,5 / 67
A. kotschoubeyanus18,9 / 5319 / 124
A. retusus3,8 / 264,3 / 69
A. scaphirostrus0 / 372 / 50
A. trigonus18,5 / 275,1 / 39

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Habitat loss and the emergence of new areas

Change in area judged suitable by the BRT model. All values refer to the baseline area of habitats assessed as suitable. Net loss is the loss minus the newly suitable area. [Breslin et al. 2020, p. 13217, corrected Table 1]
Horizon and RCPLoss of current areaNewly suitable areaNet loss
2050 · 2.646%19%27%
2050 · 4.548%10%38%
2050 · 6.047%14%33%
2050 · 8.552%25%27%
2070 · 2.650%29%21%
2070 · 4.550%5%45%
2070 · 6.050%6%44%
2070 · 8.555%2%53%

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Characters useful for identification

Selected characters from the results and key of the 2019 revision. Areole dimensions were converted from centimetres to millimetres. Ranges apply to the material and treatment of this study. For E. pachyrhiza, former subsp. parvula was examined directly. [Aquino et al. 2019, pp. 606, 608, 610–612, 615]
SpeciesAreoles and spinesFlower or underground part
Epithelantha bokei60–80 spines, persistent small projections on the spinesProjections visible by electron microscopy
E. crypticaOval areoles, 0,60–0,75 mm wideFlowers 11–12 mm long
E. greggii19–30 spines, round areoles, 1,15–1,50 mm wideOrdinary slender roots
E. ilariaeFewer than 40 spines, without persistent projectionsFlowers 17–18 mm long, opening in the evening
E. micromerisNarrow areoles, 0,20–0,30 mm wideFlowers 6–7 mm long
E. pachyrhizaWhite spines, elliptical areoles, 0,45–0,55 mm wideThickened roots
E. polycephalaBrown spines, areoles 0,90–1,80 mm wideThickened roots
E. potosina39–43 spines, areoles 0,50–0,70 mm longThickened roots
E. pulchra40–55 spines, round areolesThickened roots
E. spinosior19–25 spines, elliptical areolesSlender roots, part of the stem underground

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Self-pollen and foreign pollen in five treatments

Fruit set in the studied Thelocactus leucacanthus population in Querétaro[Delgado-Ramírez et al. 2021, p. 798, Table 3]
TreatmentFlowers / plantsFruitsSeeds per fruit, mean ± SD
Freely accessible flowers30 / 1030, or 100%65,76 ± 25,42
Hand pollination with foreign pollen29 / 1029, or 100%61,52 ± 45,19
Hand self-pollination24 / 80No fruits formed
Spontaneous self-pollination under cover15 / 50No fruits formed
Apomixis trial6 / 30No fruits formed

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Nine species and different dominant plants

Models of nine Thelocactus species. Data from Mosco's tables 1–2[Mosco 2019, p. 6, Tables 1–2]
SpeciesSitesLevins' niche breadthHighest permutation importance
T. bicolor2130,282Isothermality, 39,0%
T. buekii500,030Precipitation of the wettest quarter, 36,6%
T. conothelos1190,025Precipitation of the driest quarter, 61,2%
T. hastifer110,178Isothermality, 77,8%
T. hexaedrophorus1950,099Isothermality, 66,0%
T. leucacanthus730,050Isothermality, 66,0%
T. multicephalus130,100Isothermality, 33,0%
T. rinconensis470,062Precipitation of the wettest quarter, 58,4%
T. tulensis1170,020Isothermality, 54,4%

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Comparison of 132 adult plants

Number of complete ribs in plants studied by González-Zamora and coauthors[González-Zamora et al. 2023, p. 95, Table 5]
GroupNumber of plantsMean ± SDRange
S. coptonogonus5313,0 ± 0,3912–14
S. obvallatus (S. pentacanthus in the study)5637,0 ± 4,930–53
S. × irregularis2319,0 ± 3,615–28

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Two-year-old plants during summer warming

Microclimate of the summer experiment. Means ± standard error[Aragón-Gastélum et al. 2017b, p. 95, microclimate results]
ParameterControl plotWarming chambers
Mean air temperature21,5 ± 0,3°C23,4 ± 0,4°C
Mean daily maximum33,5 ± 0,7°C39,3 ± 1,0°C
Mean daily minimum12,5 ± 0,4°C12,8 ± 0,4°C
Relative air humidity69,6 ± 2,1%65,9 ± 2,4%

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Three Neoraimondia solution concentrations

Water parameters after application of Neoraimondia arequipensis extract. Means ± SD[Choque-Quispe et al. 2020, p. 92, Table 1]
Extract solutionTurbidity removalFlocculation indexpHBOD, mg O₂/l
1%92,64 ± 0,33%28,41 ± 0,946,75 ± 0,033,08 ± 0,02
2%92,79 ± 0,18%30,36 ± 0,116,93 ± 0,043,32 ± 0,02
3%92,78 ± 0,09%30,38 ± 0,117,02 ± 0,013,34 ± 0,03

Context and sources in the article

Structure helps identify the plant

Examples of families and water-storage organs
FamilyExampleStorage organ
CactaceaeCarnegiea giganteaStem
AizoaceaeLithops ruschiorumLeaf
CrassulaceaeCrassula deceptorLeaf
AsphodelaceaeAloe striataLeaf
AsparagaceaeAgave shawiiLeaf
ApocynaceaeHoodia gordoniiStem

Context and sources in the article

Data from studies and collections

26 results and parameters with measurement conditions ↗

Rootstocks

Separate comparison of 19 grafting rootstocks ↗