Cacti / Tabele
Tabele Botanical terms and comparisons Comparisons from the articles gathered in one place. On a narrow screen, the table can be scrolled horizontally.
Structural elements and how to recognize them Table 1 • Growth habit and general plant appearance Table 2 • Stem, branches and segments Table 3 • Spines, bristles and hairs Table 4 • Shoot surface, leaves, roots and fruits Table 5 • The flower and its parts Examples of updates using POWO, checked 9.10.2026 PTMK membership in historical tabulations Osmotic pressure of photosynthetic parenchyma after two temperature regimes (MPa) Seasonal change in segment tolerance from Saskatchewan LT₅₀ after acclimatization and improvement relative to 25/15°C LT₅₀ of organs of Opuntia ficus-indica clone 1279 (°C) Tissue LT₅₀ after 100 days at 10/5°C (°C) Damage to three-year-old opuntia segments after frost in Mendoza (%) Physiological symptom and checking its cause Document status on 9.10.2026. Use depends on the full label Products still registered. Current scope and restrictions Historical names whose old recipes must be rejected Tubercle traits and botanical examples Angle of the stem axis to the horizontal in the 1980 study <span data-cite="G:AN12" data-locator="tabela 1. SD oznacza odchylenie standardowe"></span> Categories according to the Results section of the 2024 publication <span data-cite="G:AN13" data-locator="s. 6–7"></span> Comparison of growth forms and flowers <span data-cite="G:AN03" data-locator=""></span><span data-cite="G:AN05" data-locator=""></span> Selected chromosome count results in the 2019 paper <span data-cite="G:SC06" data-locator=""></span> <a class="literature-citation web-citation" href="https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:138720-1" target="_blank" rel="noopener" title="Plants of the World Online">[2-WWW]</a> Successive runoff episodes. Distances as mean ± standard deviation <span data-cite="G:SC09" data-locator=""></span> Assembled sequences of two isolates <span data-cite="G:SC17" data-locator=""></span> <span data-cite="G:SC18" data-locator=""></span> Daily components of leaf CO₂ uptake. Conditions 25/15°C, day/night of 12 h each <span data-cite="G:CP01" data-locator=""></span> Germination after 30 days. Five replicates of 25 seeds per treatment <span data-cite="G:CP07" data-locator=""></span> Measures of ant–cactus association <span data-cite="G:NP11" data-locator=""></span><span data-cite="G:NP26" data-locator=""></span> Stem-segment junction tension. Medians in the experiment by Mylo and colleagues (2022) <span data-cite="G:NP17" data-locator=""></span> Course of healing under the experimental conditions of 30/20°C <span data-cite="G:NP19" data-locator=""></span> Levels of wood organisation in Cylindropuntia imbricata <span data-cite="G:NP21" data-locator=""></span> Correctly detected <em>S. wrightiae</em> relative to 480 plants in the ground census. Data from Bates et al. (2022), Table 2. Percentages were calculated from total counts. <span data-cite="G:PKS02" data-locator="tab. 2, s. 7"></span> IC<sub>50</sub> concentrations in enzyme tests, in µg/mL. Mean ± SD, three replicates described by the authors as biologically independent. A dash indicates the absence of a value in the source table. <span data-cite="G:PKB01" data-locator="tab. 2, s. 8"></span> Daytime microclimate beside <em>Eulychnia breviflora</em>. Means from four sensor pairs, Table 1 of Stanton and colleagues. <span data-cite="G:K2E02" data-locator=""></span> Germination of surviving <em>Eulychnia acida</em> seeds in the experiment by Cares and colleagues. 100 seeds per group, about 26°C, photoperiod 14:10 h. <span data-cite="G:K2E03" data-locator=""></span> Germination of seeds from the Santa Cruz population after four months. Mean ± standard error, three replicates of one hundred seeds each. <span data-cite="G:K2J05" data-locator="metody na s. 22 i tab. 2 na s. 24"></span> Traits of the lineages compared in the 2019 studies. <span data-cite="G:K2T01" data-locator="Vázquez-Sánchez i wsp., s. 409, 412, 416–417. De la Rosa-Tilapa i wsp., wersja online, s. 2–4"></span> <span data-cite="G:K2T02" data-locator="Vázquez-Sánchez i wsp., s. 409, 412, 416–417. De la Rosa-Tilapa i wsp., wersja online, s. 2–4"></span> Site elevations in the data of Donati and colleagues. Ranges concern the dataset from the 2017 paper. <span data-cite="G:K2T09" data-locator="wersja online, s. 5–7"></span> Fibre-wall thickness in selected radial spines, in µm. Mean ± standard deviation. <span data-cite="G:K2T03" data-locator="tab. 2, s. 122–123"></span> Shoot number and explants with hyperhydricity symptoms in <em>T. valdezianus</em>. Comparison from Table 1 of the publication, described as an assessment after 12 weeks. <span data-cite="G:K2T04" data-locator="tab. 1, s. 178"></span> Abundance over a total area of 490 m². Data from Table 2 of the 2026 study. <span data-cite="G:LOE01" data-locator="tab. 2, s. 5"></span> Diagnostic features of the underground stem and root of <em>Lophophora williamsii</em>. Histological study of five plants. <span data-cite="G:PH03" data-locator="s. 582–589, tab. 2"></span> Random transects in six known populations of <em>Lophophora williamsii</em>. Each transect occupied 100 m². Data from May and July 2019, from Table 6 of the publication. <span data-cite="G:LOW04" data-locator="s. 156, tab. 6"></span> Seedling growth after six months of the experiment. Both light levels combined. Mean ± standard deviation. <span data-cite="G:PH04" data-locator="tabela 4, s. 1770"></span> Response of Obregonia to additives in the common CM medium. Mean ± standard deviation. Data pooled for two explant types. <span data-cite="G:LOB01" data-locator="tabela 3, s. 34"></span> Flowering and recruitment in one population in Querétaro. The initial census included 468 plants in an area of approximately 20 × 30 m. Searches for new seedlings were conducted twice a year. <span data-cite="G:LOR01" data-locator="tab. 1"></span> Comparison of seed batches in the experiment conducted in 2018. Each batch comprised 50 seeds in ten dishes. Germination means the observed emergence of the radicle, and survival concerns established seedlings at the end of the short laboratory observation. The last column gives the mean ± standard error. <span data-cite="G:LOR02" data-locator="s. 5–7, ryc. 3–4"></span> Comparative characters of the taxa described. Botanical ranges from Arias and Sánchez-Martínez (2010), table 1. <span data-cite="G:P1S03" data-locator="s. 621, tabela 1"></span> Return to the surface after controlled submersion to a depth of 8 cm. 40 different seeds per taxon, mean ± standard deviation. <span data-cite="G:P1S01" data-locator="s. 91, 94"></span> Germination at 25°C with 12 hours of light, on moist filter paper. Eight replicates of 30 seeds per taxon, observed for 30 days. Mean ± standard deviation. <span data-cite="G:P1S01" data-locator="s. 91, 94"></span> Tissue structure and refractometric nectar reading. Nectar collected at 15 hours in February 2018 from 15 flowers of different individuals of each taxon in the UNAM collection. The publication does not unambiguously define the measure of variation following the ± sign. <span data-cite="G:P1S02" data-locator="s. 102–104"></span> Process elasticities in the P. bradyi model. Values from table 2 of the 2026 paper. Scopes of three studies. Demographic data and habitat comparisons represent separate parts of the analyses. Selected multiplication results from 2002. Number of shoots per initial explant, mean ± standard error. <span data-cite="G:P1P01" data-locator="tab. 1–2, s. 74–75"></span> Micrografting of <em>P. aselliformis</em> onto <em>O. ficus-indica</em>. Mean ± standard error in the 2016 experiment. <span data-cite="G:P1P02" data-locator="s. 3–4, tab. 1"></span> Flowering of Neolloydia conoidea in Tolimán. Weekly observations. Mean fruit set ± standard error, the proportion of flowers forming fruits. Data from Arroyo-Pérez et al. 2019, table 1 and sample description. <span data-cite="G:P2N02" data-locator="s. 581–583, tabela 1"></span> Controlled pollination of Neolloydia conoidea in Tolimán. 20 flowers per treatment. Fruit set is the percentage of flowers forming fruits. Seed set is the mean proportion of ovules forming seeds according to table 3 of Arroyo-Pérez et al. 2021. <span data-cite="G:P2N03" data-locator="s. 32, 35, tabela 3"></span> Multiplication of Ortegocactus macdougallii after 110 days, with NAA 4,4 µM in every treatment. Means ± standard error. Eight vessels containing six explants each per treatment, with the experiment repeated twice. Data from Arellano-Perusquía et al. 2013, table 2. <span data-cite="G:P2O01" data-locator="s. 161, 163, tabela 2"></span> Rooting of Ortegocactus macdougallii shoots after 50 days. Means ± standard error. Eight vessels containing three explants each per treatment. IBA concentrations are given in µM in accordance with the methods and results descriptions. Data from Arellano-Perusquía et al. 2013, table 4. <span data-cite="G:P2O01" data-locator="s. 161, 164, tabela 4"></span> Potential habitat area in three models. Values from table 2 of Solórzano et al. (2026). Oehmea. Atlas classification and combinations from 2026. POWO status checked on 11 October 2026. Meristem dome width in eight stages of the San Ángel material studied. Mean ± standard deviation, from table 2 of López-Ruiz et al. (2026). <span data-cite="G:NFDEV01" data-locator="s. 1333, tabela 2. cechy stadiów, s. 1328, 1330–1334"></span> Spatial associations in the study by Peters et al. (2008). “Positive” means more plants than expected in a given microhabitat, and “negative” means fewer. Accepted names follow POWO. Historical supertexta groups are retained separately. <span data-cite="G:NFDEV15" data-locator="s. 596, tabela 1"></span> <span data-cite="G:NFDEV16" data-locator="synonimy M. supertexta, 11 października 2026"></span> <span data-cite="G:NFDEV04" data-locator="wykaz akceptowanych gatunków, 11 października 2026"></span> Serotiny in Mammillaria pectinifera. Estimates for 238 m² in Texcala Snowcap in vitro. Medium stages according to table 1 and the results of the 2023 paper. Snowcap. Flower initiation in vitro after plant preparation and paclobutrazol treatment Selected site associations with soil and landscape map categories. Results of the frequency analysis in Aquino and co-authors (2021). <span data-cite="G:NFMIN01" data-locator="s. 4524–4525, ryc. 3"></span> Selected biomineral forms and examples directly shown. Plant names follow the current atlas treatment. <span data-cite="G:NFMIN03" data-locator="s. 3–7, ryc. 2, 4–6"></span> Chemical identification of selected seed samples. The scope of the results corresponds to the material identifications in Chen and co-authors (2013). <span data-cite="G:NFMIN04" data-locator="s. 203–208, ryc. 4–5"></span> Selected mean spine cell-wall thicknesses in the study by Řepka and Gebauer (2012). Measurements were made on hydrated preparations. Unit µm. <span data-cite="G:NFSP01" data-locator="s. 141, tabela 2"></span> Germination under the experimental conditions. Age means the estimated age of the tubercle zone, and the percentage is the mean reported in the publication. <span data-cite="AR:R02" data-locator="s. 364–365, tabele 1–2"></span> Response to hand self-pollination. Microscopy and fruit set were assessed in separate flowers. n denotes the size of each sample. <span data-cite="AR:R06" data-locator="s. 535, tabela 1"></span> Change in area judged suitable by the BRT model. All values refer to the baseline area of habitats assessed as suitable. Net loss is the loss minus the newly suitable area. <span data-cite="G:UCO01" data-locator="s. 13217, poprawiona tabela 1"></span> Selected characters from the results and key of the 2019 revision. Areole dimensions were converted from centimetres to millimetres. Ranges apply to the material and treatment of this study. For <em>E. pachyrhiza</em>, former subsp. <em>parvula</em> was examined directly. <span data-cite="G:UCO04" data-locator="s. 606, 608, 610–612, 615"></span> Fruit set in the studied Thelocactus leucacanthus population in Querétaro<span data-cite="G:UTHE06" data-locator="s. 798, tabela 3"></span> Models of nine Thelocactus species. Data from Mosco's tables 1–2<span data-cite="G:UTHE05" data-locator="s. 6, tabele 1–2"></span> Number of complete ribs in plants studied by González-Zamora and coauthors<span data-cite="G:UST01" data-locator="s. 95, tabela 5"></span> Microclimate of the summer experiment. Means ± standard error<span data-cite="G:PKE11" data-locator="s. 95, wyniki mikroklimatu"></span> Water parameters after application of Neoraimondia arequipensis extract. Means ± SD<span data-cite="G:UNE03" data-locator="s. 92, tabela 1"></span> Examples of families and water-storage organs
Names of cactus parts Structural elements and how to recognize them Element Meaning How to interpret it Shoot The stem together with leaves and buds, even when leaves are highly reduced The green body does not include roots. Shoot and stem are sometimes used as shorthand, but are not fully equivalent. Growing tip The region containing the shoot apical meristem New sections of the axis and organ primordia form here. Not every areole is a growing tip. Areole A specialized short shoot with its own meristematic region It can produce spines, hairs, lateral shoots or flowers. It is not a single spine or a hole in the epidermis. Rib A longitudinal ridge on the stem Its geometry facilitates volume changes in the hydrated shoot. Rib number may change during growth. Tubercle A distinct projection on the shoot surface It may be associated with an areole. Its arrangement and grooves matter in genus descriptions. Axil In mammillaria descriptions, the depression between tubercles The flowering region may be separate from the spine-bearing apical portion of the areole. Radial spine A spine arranged radially near the areole's perimeter It usually lies closer to the shoot surface. The distinction from a central spine concerns position, not just length. Central spine A spine developing nearer the centre of the spine cluster It often projects farther from the surface. It may be straight or hooked. Glochid A small barbed spine typical of opuntioids It detaches easily and penetrates skin. Glochids are not soft hairs that can be touched safely. Areole hairs Trichomes, often forming a woolly covering They should be distinguished from thin or feathery spines. Similar appearance does not demonstrate shared structure. Segment A shoot section separated by a distinct constriction A flat opuntia or schlumbergera segment is part of the stem, not a leaf. Epidermis and cuticle The covering tissue and its outer protective layer Stomata belong to the epidermis. The cuticle limits water loss but does not seal the plant completely. Photosynthetic parenchyma Living cells with chloroplasts It performs photosynthesis. It usually lies in the outer portion of the shoot cortex. Water-storage tissues Living tissues maintaining a water reserve They are not an empty cistern. Storage and photosynthetic functions may partly overlap. Xylem and phloem Conducting tissues Xylem carries water and minerals, while phloem mainly carries organic compounds between sites of production and use. Root collar The transition zone between shoot and root system Its position helps assess correct planting depth and the location of damage. Primary root and lateral roots The axis of embryonic origin and its branches Roots arising from the shoot of a cutting are adventitious. A large root is not necessarily a primary root. Storage root A thickened root storing resources It is not an underground stem. Cacti differ in the contribution of roots and shoots to water storage.
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Growth habit and general plant appearance Table 1 • Growth habit and general plant appearance Element or term Meaning Examples and explanation phyton Plant, an element of Greek origin Astrophytum combines a reference to a star with the term for a plant. Conophytum is an example outside Cactaceae.stēlē, columna Column or pillar chrysostele , albicolumna . stēlē comes from Greek, columna from Latin.dendron, arbor Tree arborescens means becoming tree-like. The element dendro- is Greek.dumus Shrub or scrub dumetorum refers to scrub as a habitat and need not describe the cactus's own habit.frutex Shrub fruticosus , frutescens indicate a shrubby character or its development.virgultum Shrub, scrub, young shoots virgultus should be read in the context of the original description.sphaira, globus Sphere sphaerocarpus applies globose shape to a fruit, globularis to a spherical shape. Not every element refers to the entire plant.clava, korynē, rhopalon Club clavatus , claviformis , corynodes . They describe broadening towards the end of an organ.cylindrus, kylindros Cylinder cylindroarticulatus means cylindrically segmented. The element also occurs in Cylindropuntia .discus Disc, shield disciformis means disc-shaped. Discocactus illustrates the element's use in a genus name.caespes Turf, dense clump caespitosus means clumping, forming clusters of shoots.depressus Flattened, lowered Describes organ proportions. A flattened stem need not be segmented. prostratus, pendulus Prostrate, hanging Two distinct growth directions. They should not be treated as synonyms.
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Stem, branches and segments Table 2 • Stem, branches and segments Element or term Meaning Examples and explanation caulis, kaulos Stem Latin caulis and Greek kaulos denote the shoot axis. fusicaulis refers to a spindle-shaped stem. truncus Trunk The correct Latin term for a trunk. The former assignment of “trunk” to stemma was incorrect. ramus Branch ramosus means branched, ramosissimus very strongly branched.brachium Arm brachiatus describes arm-like branches.klados Branch, offshoot heterocladus refers to differences between branches, glaucocladus to their glaucous colour.frons Leafy branch or blade, depending on context latifrons and acutifrons require checking the organ description. The element alone does not demonstrate typical leaves.sarmentum Long, slender shoot sarmentaceus is associated with this habit. The archive used the inaccurate spelling sarmentium .articulus, arthron Segment or joint articulatus means segmented. Arthrocereus contains a Greek element referring to joints between segments.stemma Wreath, crown This Greek element remains in the comparison as a correction. It is not a term for a stem or trunk.
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Spines, bristles and hairs Table 3 • Spines, bristles and hairs Element or term Meaning Examples and explanation spina, akantha Spine, sharp structure tenuispinus , melanacanthus . Etymology does not replace identification of an organ's origin.aculeus Prickle or sharp outgrowths in older descriptions aculeatus . In a strict morphological description, a prickle is a surface outgrowth, while a cactus spine has leaf origin.ankistron, hamus, uncus Hook ancistrophorus , multihamatus , uncinatus indicate a hooked curvature.centrum Centre centrispinus refers to central spines. Position in the areole is distinct from spine length.acutus, acumen Sharp, sharply pointed tip acuminatus indicates a gradually elongated point. The imprecise archival spelling aculius was omitted.cuspis Point, sharp ending tricuspidatus means three-pointed. The number refers to a particular feature in the description.subula Awl subulatus describes an awl-shaped form tapering to a sharp end.acus Needle acifer , acispinosus refer to a needle-like form.arma Armament Armatocereus , albiarmatus . A metaphor for defensive traits.acinaces Short curved sword acinaciformis , acinacispinus describe a shape resembling a blade.machaira Blade, knife or sword Machaerocereus and machaeratus illustrate historical use of the element, not a statement of current genus status.pugio Dagger pugionacanthus , pugionifer refer to the shape or presence of dagger-like structures.aor, ensis, gladius, xiphos Sword ensiformis , gladiispinus , xiphacanthus . These are different roots evoking similar shapes.hasta, lancea Spear, lance hastatus and lanceolatus are not interchangeable. In leaf descriptions they denote different outlines.sarissa Long spear sarrisophorus refers to long, sharp armament.chaitē, seta Bristle chaetacanthus , saetiger , acanthosetus emphasize a thin, bristly form.pilus, thrix Hair Latin pilo- and Greek tricho- . Examples include Pilosocereus and sphaerotrichus . coma, crinis Tuft, hair albicomus , brevicrinita . A reference to hair does not establish whether the structure described is a trichome or a thin spine.erion, lana Wool eriophorus , laniceps indicate a woolly appearance of a particular part.floccus, cincinnus Tuft, curl flocculosus , cincinnatus describe covering arrangement, not a separate cactus organ.pinna, pluma Feather pinnatus , plumosus . In cacti, a feathery appearance may result from spine structure.inermis Unarmed Closest in meaning to lacking spines or prickles. subinermis means almost unarmed. nudus Naked, exposed nudiflorus applies nakedness to the flower. It does not automatically mean the entire plant lacks spines.calvus Bald, without covering calvescens indicates becoming bald, losing the covering during development.laevis Smooth laevigatus refers to a smooth surface. Smoothness and lack of spines are different traits.glaber Hairless glabrescens means becoming hairless. It concerns hairs, not a general statement of spinelessness.
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Shoot surface, leaves, roots and fruits Table 4 • Shoot surface, leaves, roots and fruits Element or term Meaning Examples and explanation areola Small area, in cacti an areole longiareolatus refers to elongated areoles. An areole is the site where a specialized short shoot develops.glochis Barb, barbed point glochidiatus refers to glochids. The singular in a Polish description may be rendered as glochidium.derma Skin Argyroderma is an example outside Cactaceae. A surface name does not establish the anatomical covering type.sperma, semen Seed cardiospermus , dolichospermaticus . The elements may describe shape or other seed traits.fructus, karpos Fruit albifructus , platycarpus apply a trait to the fruit.bacca Berry baccifer means producing berries. It is not a synonym for every fruit type.pulpa Pulp duripulpa refers to pulp hardness.radix, rhiza Root brunneiradicatus , macrorhizus , polyrhizus . The term may concern root colour, size or number.rapum, napus Turnip napiformis describes resemblance to a turnip, usually organ thickening. It is not a general term for a root.angulus, gonia Angle multangularis , hexagonus , heptagonus . An angular cross-section may correspond to ribbing, but “angle” and “rib” are not exact synonyms.costa, pleura Rib, side paucicostatus means having few ribs, while pleurocarpus applies ribbing or lateral position to the fruit, depending on the name.mamma, mastos Breast, projection grandimamma , Echinomastus refer to tubercles or resemblance in shape.papilla, thēlē Papilla, tubercle papillosus , conothele , Thelocactus . Not every microscopic surface papilla is a large shoot tubercle.folium, phyllon Leaf grandifolius , rotundifolius , phyllacanthus . The former name Phyllocactus referred to a leaf-like stem, not ordinary leaves.
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The flower and its parts Table 5 • The flower and its parts Element or term Meaning Examples and explanation anthos, flos Flower micranthus , grandiflorus . The first element is Greek, the second Latin.kalyx, calyx Calyx Gymnocalycium refers to a “naked calyx”. The historical meaning does not replace a modern description of the floral tube and pericarpel.sepalum Sepal crassisepalus , smaragdisepalus . In many cacti, the boundary between outer and inner tepals is gradual.petalon, petalum Petal oxypetalus refers to sharply pointed petals. In a description of a continuous perianth, tepals may be the safer term.stylos, stylus Style, originally a pillar or stylus brevistylus , flavistylus describe style length or colour.graphis Stylus, writing tool The old brachygraphis illustrates a shape metaphor. graphis should not be used as a universal term for the style. stamen, stēmōn Stamen longistamineus refers to stamens. A stamen includes a filament and anther, so it is not the filament alone.filamentum Thread, including a stamen filament rubrifilamentosus indicates red filaments. filamenta is plural.anthera, stigma, ovarium Anther, stigma, ovary Three supplementary anatomical terms. The anther produces pollen, the stigma receives it, and the ovary contains ovules.
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Former names in the supplied books Examples of updates using POWO, checked 9.10.2026 Former name Accepted name Relationship and significance Zygocactus truncatus Schlumbergera truncata Changed generic combination. We retain the former name when discussing the book. [2-WWW] Chamaecereus silvestrii Echinopsis chamaecereus The accepted name follows current POWO. The older combination helps find the earlier description. [2-WWW] Mammillaria wildii Mammillaria crinita subsp. wildii It has subspecies rank in the adopted treatment. [2-WWW] Hylocereus undatus Selenicereus undatus Transfer to another genus does not mean a different species cultivated as a rootstock. [2-WWW] Dolichothele longimamma Mammillaria longimamma A former genus found in literature describing elongated tubercles. [2-WWW] Neomammillaria gaumeri Mammillaria heyderi subsp. gaumeri The name from Britton and Rose's monograph now corresponds to a subspecies in POWO's treatment. [2-WWW]
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Exhibitions and the scale of development PTMK membership in historical tabulations Date of tabulation Members Data scope 18 November 1966 388 Total excluding Bydgoszcz data. Includes 25 people from provinces where branches had not been constituted. Second General Congress, June 1969 1198 13 branches. The authors separately record 15 corresponding members from other countries. Third General Congress, 1972 About 300 10 branches. The source gives an approximate value.
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Water loss, sugars and mannitol Osmotic pressure of photosynthetic parenchyma after two temperature regimes (MPa) Material identified in the study 30/20°C, day/night 10/0°C, day/night Opuntia ficus-indica 0.70 ± 0.02 0.80 ± 0.01 Opuntia humifusa 0.88 ± 0.05 1.23 ± 0.06 Opuntia streptacantha 0.71 ± 0.02 0.83 ± 0.03
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Seasonal resistance on the Canadian prairie Seasonal change in segment tolerance from Saskatchewan Date and material Water (% fresh mass) LT₀, no visible damage LT₅₀, tissue damage 6 IX 1985, mature segments O. fragilis 84.3% −7°C −10°C 10 X 1985, O. fragilis 71.9% −50°C below −70°C 10 XII 1985, O. fragilis 69.6% −50°C below −70°C 8 IV 1986, O. fragilis 72.5% −30°C below −60°C 18 VI 1986, young segments O. fragilis 91.0% not determined death after freezing at −3.5°C 10 XII 1985, O. polyacantha 67.6% −40°C −55°C
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Twenty populations of Opuntia fragilis LT₅₀ after acclimatization and improvement relative to 25/15°C Material provenance LT₅₀ (°C) Tolerance improvement (°C) Peace River, Alberta −39.0 20.0 Fort St. John, British Columbia −44.3 24.3 Goose Lake, Saskatchewan −41.1 23.4 Cache Creek, British Columbia −34.9 14.9 Bird River, Manitoba −22.6 17.2 Vernon, British Columbia −30.1 11.1 Hornby Island, British Columbia −19.1 12.1 Lethbridge, Alberta −28.3 19.0 Virden, Manitoba −48.9 19.9 Minot, North Dakota −40.2 17.2 Plymouth, Washington −20.6 11.1 Eau Claire, Wisconsin −25.5 18.8 Kaladar, Ontario −20.4 15.4 Thermopolis, Wyoming −28.6 17.6 Salt Lake City, Utah −25.0 17.0 Leadville, Colorado −22.4 16.4 Princeton, Colorado −15.6 14.2 Russell, Kansas −23.3 19.3 Loa, Utah −23.6 13.6 Albuquerque, New Mexico −27.6 19.6
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Shoots, roots, fruits and meristems respond differently LT₅₀ of organs of Opuntia ficus-indica clone 1279 (°C) Organ and age After 20/10°C After 30/20°C One-year segment −7.6 ± 0.2 −6.7 ± 0.2 Six-week segment −7.8 ± 0.2 −6.5 ± 0.1 Six-week fruit −6.3 ± 0.3 −6.3 ± 0.2 Six-week root −6.5 ± 0.2 −6.4 ± 0.2
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Shoots, roots, fruits and meristems respond differently Tissue LT₅₀ after 100 days at 10/5°C (°C) Material in the publication Photosynthetic parenchyma Areole meristem Nopalea cochenillifera −9.1 ± 0.4 −11.3 ± 0.4 Opuntia robusta −8.8 ± 0.4 −11.2 ± 0.4 Selenicereus megalanthus −6.5 ± 0.3 −7.4 ± 0.4
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Field studies and a real winter Damage to three-year-old opuntia segments after frost in Mendoza (%) Material named in the publication Plain Dune Mean O. paraguayensis 0.0 0.7 0.3 O. robusta 3.0 0.9 2.0 O. spinulifera 3.7 6.5 5.1 O. ficus-indica “San Rafael”19.2 10.6 14.9 O. ficus-indica “Cuenca”25.7 23.6 24.6 O. ficus-indica “San Juan”52.6 16.1 34.4
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Differential diagnosis table Physiological symptom and checking its cause Observation What to check Course of action Pale patch after a change of position Relationship to exposure, side of the stem, temperature Gradual acclimatisation and observation of new growth Narrow, elongated growth Typical growth form and lighting history Improved light suited to the growth stage Wilting in a wet pot Root condition, drainage, duration of moisture retention Restoring suitable root conditions and distinguishing infection Chlorosis after intensive fertilisation EC, pH and root viability Identifying the problem before further fertilisation Watery tissue after cooling Temperature history and expansion of damage Isolating the plant and assessing lasting damage and secondary rot
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Products checked in official documents Document status on 9.10.2026. Use depends on the full label Product and user What the document states Relevance to cactus cultivation Sanium AL Non-professional Flupyradifurone, a ready-to-use spray product. Ornamental plants indoors and under cover, against pests including aphids, scale insects and greenhouse whitefly. [10-WWW] Do not extend this scope to root drenching. Outdoor uses differ. The full label applies. Mospilan 20 SP Professional Acetamiprid. The label dated 11.08.2025 includes specified minor uses in ornamental plants under cover and in the ground. [10-WWW] A product for professional users. The spraying scope does not include arbitrary drenching of the root ball. Magnicur Energy Non-professional Propamocarb and fosetyl. The label dated 24.02.2026 includes Phytophthora disease and diseases caused by Pythium in ornamentals, under specified conditions and at specified stages. [10-WWW] It is not a universal product for every rot. The cultivation site, plant stage and diagnosis must be checked for the particular method. Actellic 500 EC Professional Pirimiphos-methyl. The current label concerns disinsection of warehouses and protection of specified cereal grain. [10-WWW] The historical recommendation to use it against cactus pests does not match the current scope of this label. Kaptan Gold 80 WG R-101/2021 The register gives 30.09.2026 as the deadline for using existing stocks. [10-WWW] The deadline had passed by the verification date. This does not mean that all other captan-containing products have been withdrawn.
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Historical names still found in the current register Products still registered. Current scope and restrictions Product and user Current use Comment on the historical book Basamid Professional. Fumigation training required Dazomet. Soil or substrate disinfection before planting and sowing, within the scope specified by the label. [10-WWW] The product remains in the register. This is a specialist substrate preparation treatment, rather than treatment of plants in a household pot. Siarkol 80 WP Professional Sulphur, a fungicide. The label includes powdery mildew of ornamental plants. [10-WWW] The current formulation and its use are given. Historical sulphur use against “red spider” has not been carried over to the scope of this label. Kaptan Zawiesinowy 50 WP Professional Captan. The label dated 7.11.2025 includes apple and pear scab and, in permanent greenhouses, scab and leaf spots of ornamental trees and shrubs. For annual ornamental plants, it includes grey mould. [10-WWW] Do not equate it with Kaptan Gold 80 WG or an old general recipe for cacti. The range of crops and diseases listed on this label applies. Miedzian 50 WP R-134/2015 Professional and non-professional uses Copper oxychloride. Uses include apple scab and fire blight. The documents specify the scope and method of use separately for professionals and amateurs. [10-WWW] The brand remains in the register. Orchard and vegetable uses are not automatically extended to control of cactus rots.
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Historical products not adopted in the recommendations Historical names whose old recipes must be rejected Names in historical guides Comparison result Conclusion Benlate, Fundazol, Dithane M-45, Novozir MN 80 No current products under these exact names were found in the register checked. [10-WWW] Do not reproduce historical concentrations or mixtures. Any new product is assessed according to its own authorisation. Ekatin, Anthio, Metasystox R, Rogor, Ultracid 40 EC These exact names are absent from the list checked. [10-WWW] Historical mealybug or mite control is not a current application instruction. DDT and mercury products described historically They have not been adopted in the recommendations. Fleischer and Schütz already noted restrictions on older products in their 1978 edition. [Fleischer and Schütz 1978, pp. 188–193] The history of cultivation advice is retained without reproducing obsolete practice. General names “sulphur”, “copper”, “captan” The substance name alone does not specify a product or its current scope. [10-WWW] Do not label an entire substance as authorised or withdrawn on the basis of one brand.
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How to compare the eight diagrams Tubercle traits and botanical examples Trait compared Example Basis of interpretation Short-conical Mammillaria mammillaris Morphological description [Britton and Rose 1923, p. 70] , current name [2-WWW] Elongated Mammillaria longimamma Morphological description [Britton and Rose 1923, pp. 61–63] , current name [2-WWW] Cylindrical and rounded Mammillaria bocasana Morphological description [Britton and Rose 1923, p. 147] , current name [2-WWW] Slender and strongly tapering Mammillaria longimamma , comparison of elongationMorphological description [Britton and Rose 1923, pp. 61–63] , current name [2-WWW] Blunt-tipped Mammillaria heyderi subsp. gaumeri Morphological description [Britton and Rose 1923, pp. 72–73] , current name [2-WWW] Angular, approaching pyramidal Mammillaria polyedra Morphological description [Britton and Rose 1923, pp. 102–103] , current name [2-WWW] Laterally flattened Mammillaria compressa Morphological description [Britton and Rose 1923, pp. 90–92] , current name [2-WWW] Densely arranged Mammillaria evermanniana Morphological description [Britton and Rose 1923, p. 97] , current name [2-WWW]
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What differs between close relatives Comparison of growth forms and flowers [de Vos et al. 2025] [Korotkova et al. 2010] Trait Calymmanthium Lymanbensonia Habit Branched shrubs or trees Smaller plants, erect, spreading or hanging Stem Several thin, wing-like ribs Flattened or angular, depending on species Substrate Terrestrial growth Terrestrial, rock-dwelling or epiphytic growth Flower bud Hidden inside the short-shoot covering, which opens before flowering More typical architecture, with an exposed developing perianth
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Two historical names in one current species Selected chromosome count results in the 2019 paper [Las Peñas et al. 2019] [2-WWW] Current name Identification of material in the paper 2n Number of x sets T. alexanderi T. alexanderi 22 2 T. bonnieae T. bonnieae 44 4 T. articulatus var. strobiliformis 77 7 T. articulatus var. articulatus and var. oligacanthus 88 8 T. aoracanthus T. aoracanthus 242 22 T. aoracanthus T. paediophilus 319 29
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Sixty labelled stem segments Successive runoff episodes. Distances as mean ± standard deviation [Almirón et al. 2019] Episode Rainfall and duration Mean displacement First 35 mm in 2 days 480.4 ± 210.0 m Second 100 mm in 4 days 103.0 ± 138.9 m Third 20 mm in several hours 33.2 ± 52.3 m
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The leaf's full-day balance Daily components of leaf CO₂ uptake. Conditions 25/15°C, day/night of 12 h each [Nobel and Hartsock 1987] Species Drought [days] Day [mmol m⁻²] Night [mmol m⁻²] Pereskiopsis porteri 0 108 7 Pereskiopsis porteri 7 6 72 Pereskiopsis porteri 14 1 35 Quiabentia verticillata (as Q. chacoensis )0 129 9 Quiabentia verticillata 7 24 16 Quiabentia verticillata 14 0 2 Austrocylindropuntia subulata 0 111 2 Austrocylindropuntia subulata 7 6 24 Austrocylindropuntia subulata 14 0 15
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Two species, two responses Germination after 30 days. Five replicates of 25 seeds per treatment [Jara-Peña et al. 2024] Species and origin 12 h of light per day Continuous darkness Austrocylindropuntia floccosa , Apurímac28.0% 32.8% Cumulopuntia ignescens , Arequipa36.0% 0%
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Three levels of the result Measures of ant–cactus association [Braun and Lortie 2025] [Lynch and Savage 2023] Level What was measured Meaning Visits Number and identity of ants Resource access and activity on the plant Defence Bug numbers or response to test prey Herbivore pressure or defensive behaviour Reproduction Number of developed seeds per fruit One component of reproductive outcome
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Force and strength describe different traits Stem-segment junction tension. Medians in the experiment by Mylo and colleagues (2022) [Mylo et al. 2022] Material studied Maximum force [N] Tensile strength [MPa] Cylindropuntia bigelovii 14.4 1.41 Opuntia ficus-indica, younger lateral segment 51.2 3.84 Opuntia ficus-indica, older lateral segment 153 2.70
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Wound periderm Course of healing under the experimental conditions of 30/20°C [Mylo et al. 2020] Observation stage Tissue change First days Edge curling and a lipophilic barrier at the wound surface Around the first week Visible pale layer of dead cells and development of wall changes Subsequent weeks, through day 31 Organisation of wound periderm and remodelling of tissues beneath the surface
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Directions in wood Levels of wood organisation in Cylindropuntia imbricata [Morankar et al. 2024] Structural scale Visible elements Scope of information Stem skeleton Openings and connected wood bands Geometry of the whole structure Tissues and cells Vessels, fibres, rays Direction and type of wood organisation Cell wall Layers and pits Local structure and response to pressure
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How many plants the camera detects Correctly detected S. wrightiae relative to 480 plants in the ground census. Data from Bates et al. (2022), Table 2. Percentages were calculated from total counts. [Bates et al. 2022, Table 2, p. 7] Flight height Correctly detected Proportion of ground census Missed 10 m 183 38,1% 297 15 m 89 18,5% 391 20 m 46 9,6% 434
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Inhibition of two enzymes IC50 concentrations in enzyme tests, in µg/mL. Mean ± SD, three replicates described by the authors as biologically independent. A dash indicates the absence of a value in the source table. [Heraiz et al. 2026, Table 2, p. 8] Material tested AChE BACE-1 Spines, methanolic extract 0,362 ± 0,012 0,425 ± 0,014 Stem, methanolic extract 0,603 ± 0,019 0,939 ± 0,031 A. oryzae culture, ethyl acetate extract0,255 ± 0,008 0,550 ± 0,018 A. oryzae culture, subsequent methanolic extract3,180 ± 0,104 1,882 ± 0,062 Donepezil, reference substance for AChE 0,167 ± 0,006 — Curcumin, reference substance for BACE-1 — 0,350 ± 0,012
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Cooler air beside the stem surface Daytime microclimate beside Eulychnia breviflora . Means from four sensor pairs, Table 1 of Stanton and colleagues. [Stanton et al. 2014] Parameter With epiphytes Without epiphytes Mean daytime temperature 15,1°C 17,3°C Mean of daily maximum temperatures 20,9°C 25,1°C Mean daytime VPD 0,69 kPa 1,02 kPa Mean of daily maximum VPD values 1,54 kPa 2,33 kPa
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Seeds after passage through the digestive tract Germination of surviving Eulychnia acida seeds in the experiment by Cares and colleagues. 100 seeds per group, about 26°C, photoperiod 14:10 h. [Cares et al. 2018] Seed origin Germination Meaning of the comparison Directly from fruits 52% Control group Recovered after goats 60% No demonstrated difference from the control Recovered after guanacos 13% Lower than in the control
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The same seed batch under four conditions Germination of seeds from the Santa Cruz population after four months. Mean ± standard error, three replicates of one hundred seeds each. [Jaramillo et al. 2010, methods on p. 22 and Table 2 on p. 24] Preparation Substrate in the dish Germination [%] Soaking for 24 h at 25°C Moist filter paper 52,0 ± 10,2 Soaking for 24 h at 60°C Moist filter paper 47,3 ± 8,1 No soaking Moist soil 7,3 ± 3,0 No soaking Moist filter paper 66,7 ± 8,6
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Crystals and fibres in a precisely defined location Traits of the lineages compared in the 2019 studies. [Vázquez-Sánchez et al. 2019, Vázquez-Sánchez et al., pp. 409, 412, 416–417. De la Rosa-Tilapa et al., online version, pp. 2–4] [De la Rosa-Tilapa et al. 2019, Vázquez-Sánchez et al., pp. 409, 412, 416–417. De la Rosa-Tilapa et al., online version, pp. 2–4] Genus Seed microsculpture Hypodermal crystals Fibres in cortical bundles Turbinicarpus Striate Prismatic Absent in the material studied Rapicactus Papillate Concentric druses Absent in the material studied Kadenicarpus Smooth Prismatic Present in the material studied
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Two lineages in a mountain landscape Site elevations in the data of Donati and colleagues. Ranges concern the dataset from the 2017 paper. [Donati et al. 2017, online version, pp. 5–7] Group in the study Site elevation Band with the most taxa Rapicactus 1360–3120 m above sea level 1400–1800 m above sea level Turbinicarpus as treated by the authors307–2185 m above sea level 1000–1400 m above sea level
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A papery ribbon and a corky structure Fibre-wall thickness in selected radial spines, in µm. Mean ± standard deviation. [Mosco 2009, Table 2, pp. 122–123] Taxon studied Spine description Fibres in the deeper portion Thin-walled fibres T. valdezianus Woody structure, feathery epidermis 4,1 ± 0,9 Not found T. schmiedickeanus subsp. flaviflorus (T. flaviflorus in the paper)Papery Not found 1,5 ± 0,3 T. schmiedickeanus subsp. rioverdensis (T. rioverdensis in the paper)Corky, with fissures 3,1 ± 0,9 1,0 ± 0,2
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Multiplication and symptom frequency Shoot number and explants with hyperhydricity symptoms in T. valdezianus . Comparison from Table 1 of the publication, described as an assessment after 12 weeks. [García Osuna et al. 2011, Table 1, p. 178] Additive to the kinetin medium Shoots per explant, mean Explants with symptoms Control without the tested additives 3,55 b 25% Paclobutrazol, 3,4 × 10−4 M 18,00 a 0% Prohexadione-calcium, 10−4 M 3,25 b 50% Salicylic acid, 10−4 M 1,15 b 75% Benzoic acid, 10−4 M 1,20 b 75%
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Buds remaining underground Diagnostic features of the underground stem and root of Lophophora williamsii . Histological study of five plants. [Terry and Mauseth 2006, pp. 582–589, Table 2] Tissue or structure Underground stem Root Developmental significance Outer parenchyma Cortex with xylem and phloem bundles In older portions, a zone of secondary phloem parenchyma without a cortical bundle network The similar appearance of the swollen structure has different anatomical foundations Organ centre Parenchymatous pith without xylem elements Expanded metaxylem and inner secondary xylem with scattered conducting elements Allows organ identity to be determined in section Areoles and their meristems Living remnants can persist beneath the soil surface No areoles Retained stem buds produce new branches
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Clusters among empty plots Random transects in six known populations of Lophophora williamsii . Each transect occupied 100 m². Data from May and July 2019, from Table 6 of the publication. [Ermakova et al. 2021, p. 156, Table 6] Region and site Transects with plants / all transects Number of plants Density on transects (ha⁻¹) South, site 1 4 / 27 71 262,96 South, site 2 3 / 31 73 235,48 South, site 3 1 / 26 53 203,85 West, site 4 1 / 14 25 178,57 West, site 5 1 / 18 26 144,44 West, site 6 4 / 5 46 920,00
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Changing proportions with water availability Seedling growth after six months of the experiment. Both light levels combined. Mean ± standard deviation. [Martínez-Berdeja and Valverde 2008, Table 4, p. 1770] Substrate hydration relative to the reference value Relative growth rate, g g⁻¹ day⁻¹ Root-to-shoot dry-mass ratio Ratio of relative root and shoot growth rates, K 25% 0,007 ± 0,002 1,106 ± 0,353 2,775 ± 0,895 60% 0,009 ± 0,003 0,774 ± 0,388 1,51 ± 0,256
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Medium additives and the quality of new shoots Response of Obregonia to additives in the common CM medium. Mean ± standard deviation. Data pooled for two explant types. [Cardarelli et al. 2010, Table 3, p. 34] Medium treatment Shoots per explant Explants with callus, % Spontaneous rooting, % CM 1,0 ± 0,5 — — CM + putrescine, 1 mM 2,4 ± 0,6 16,7 ± 1,1 33,3 ± 3,7 CM + AgNO₃, 10,2 µM 1,2 ± 0,3 15,0 ± 1,0 14,6 ± 1,3 CM + CPPU, 12,1 µM 5,0 ± 0,4 36,8 ± 4,2 —
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Numbers of flowers, fruits and seedlings Flowering and recruitment in one population in Querétaro. The initial census included 468 plants in an area of approximately 20 × 30 m. Searches for new seedlings were conducted twice a year. [Briseño-Sánchez et al. 2022, Table 1] Year Number of flowers Flowers forming fruits New seedlings 2015 462 44,3% 21 2016 562 22,2% 21 2017 783 26,3% 31 2018 835 44,5% 30 2019 115 62,6% 5 2020 162 25,9% 1 2021 423 32,8% 18
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Seeds stored for four years Comparison of seed batches in the experiment conducted in 2018. Each batch comprised 50 seeds in ten dishes. Germination means the observed emergence of the radicle, and survival concerns established seedlings at the end of the short laboratory observation. The last column gives the mean ± standard error. [Briseño-Sánchez et al. 2024, pp. 5–7, Fig. 3–4] Collection year and storage Seeds in the sample Observed germination Seedling survival 2018, from the year of the experiment 50 62% 61 ± 5,92% 2014, approximately four years 50 52% 58 ± 4,81%
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A seed from a rocky canyon Comparative characters of the taxa described. Botanical ranges from Arias and Sánchez-Martínez (2010), table 1. [Arias and Sánchez-Martínez 2010, p. 621, Table 1] Taxon Stem, height × diameter Spines Perianth colour S. disciformis subsp. disciformis 3–12 × 3–9 cm 1–5, flexible, shed early White to yellowish white S. disciformis subsp. esperanzae 2,5–4,5 × 4–5 cm 1–5, flexible, shed early Intense magenta S. corregidorae 18–23 × 8–12 cm 3–5, longer and persistent Yellow
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Air in the aril and coat cells Return to the surface after controlled submersion to a depth of 8 cm. 40 different seeds per taxon, mean ± standard deviation. [Camacho-Velázquez et al. 2018, pp. 91, 94] Taxon Aril Time to return to the surface S. disciformis subsp. disciformis Present 2,61 ± 0,53 s S. disciformis subsp. esperanzae Present 1,74 ± 0,65 s S. corregidorae Absent 4,72 ± 1,55 s
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Germination at a specified temperature and light regime Germination at 25°C with 12 hours of light, on moist filter paper. Eight replicates of 30 seeds per taxon, observed for 30 days. Mean ± standard deviation. [Camacho-Velázquez et al. 2018, pp. 91, 94] Taxon Aril Germination Mean germination time S. disciformis subsp. disciformis Present 97,08 ± 3,75% 5,29 ± 0,04 days S. disciformis subsp. esperanzae Present 92,08 ± 7,75% 5,27 ± 0,06 days S. corregidorae Absent 96,67 ± 4,36% 5,31 ± 0,06 days
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Measuring nectar at a specified time Tissue structure and refractometric nectar reading. Nectar collected at 15 hours in February 2018 from 15 flowers of different individuals of each taxon in the UNAM collection. The publication does not unambiguously define the measure of variation following the ± sign. [Camacho-Velázquez et al. 2019, pp. 102–104] Taxon Epidermal cells Secretory parenchyma layers Nectar reading S. disciformis subsp. disciformis Slightly convex Approximately 10–12 22,09 ± 7,42°Brix S. disciformis subsp. esperanzae Relatively flat Approximately 8–9 8,65 ± 3,98°Brix S. corregidorae Papillate convex Approximately 7–8 16,12 ± 4,48°Brix
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Which processes had the greatest effect Process elasticities in the P. bradyi model. Values from table 2 of the 2026 paper. Process Elasticity of λ Survival 0,39 Stem growth in the visible state 0,24 Root elongation and exit from burial 0,17 Growth after the buried state 0,15 Entry into the buried state −0,01
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Twenty years in one Utah plot Scopes of three studies. Demographic data and habitat comparisons represent separate parts of the analyses. Species Material and period Main observations P. bradyi 965 plants, three plots, a 31-year database Relationship of survival to diameter and transition to the buried state P. nigrispinus Four demographic plots, censuses in 2016–2019 and 2022 Loss of recorded plants, no new seedlings detected P. winkleri 107 plants in one plot, 1995–2014 Effects of trampling and weather, reproduction concentrated in some specimens
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Cytokinin, apical dominance and successive cycles Selected multiplication results from 2002. Number of shoots per initial explant, mean ± standard error. [Pérez-Molphe-Balch and Dávila-Figueroa 2002, Table 1–2, pp. 74–75] Treatment Shoots after 60 days Shoots after 180 days P. aselliformis , apex BA 4,4 µM, sucrose 30 g/L13,23 ± 1,41 122,27 ± 9,05 P. aselliformis , apex BA 8,8 µM, sucrose 30 g/L13,70 ± 1,37 128,07 ± 9,16 P. strobiliformis , disc BA 4,4 µM, sucrose 50 g/L12,37 ± 1,21 119,33 ± 9,66 P. strobiliformis , apex BA 4,4 µM, sucrose 30 g/L6,43 ± 0,40 136,33 ± 12,6
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The apex and an areole-bearing segment give different results Micrografting of P. aselliformis onto O. ficus-indica . Mean ± standard error in the 2016 experiment. [Badalamenti et al. 2016, pp. 3–4, Table 1] Measure Apical scion Subapical segment Scion length 5 mm 3 mm Number of micrografts 30 270 Successful micrografts 97 ± 0,7% 81 ± 1,2% New shoots per scion 1,0 ± 0 2,7 ± 0,3 Length after 3 weeks 31 ± 0,6 mm 28 ± 0,4 mm Length after 6 weeks 38 ± 0,3 mm 35 ± 0,5 mm
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Five or six flowering waves Flowering of Neolloydia conoidea in Tolimán. Weekly observations. Mean fruit set ± standard error, the proportion of flowers forming fruits. Data from Arroyo-Pérez et al. 2019, table 1 and sample description. [Arroyo-Pérez et al. 2019, pp. 581–583, Table 1] Year Plants in the analysis Flowering waves Recorded flowers Fruit set 2015 146 6 930 0,67 ± 0,022 2016 142 5 907 0,58 ± 0,023
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Pollen from other plants and the number of seeds formed Controlled pollination of Neolloydia conoidea in Tolimán. 20 flowers per treatment. Fruit set is the percentage of flowers forming fruits. Seed set is the mean proportion of ovules forming seeds according to table 3 of Arroyo-Pérez et al. 2021. [Arroyo-Pérez et al. 2021, pp. 32, 35, Table 3] Treatment Fruit set Seed set Insect access, control 90% 0,73 Hand cross-pollination 95% 0,96 Hand self-pollination 0% 0 Isolation without pollination 0% 0
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Multiplication, shoot numbers and length Multiplication of Ortegocactus macdougallii after 110 days, with NAA 4,4 µM in every treatment. Means ± standard error. Eight vessels containing six explants each per treatment, with the experiment repeated twice. Data from Arellano-Perusquía et al. 2013, table 2. [Arellano-Perusquía et al. 2013, pp. 161, 163, Table 2] BA (µM) Shoots per explant Shoot length (mm) Fresh mass (mg) 13,3 5,67 ± 0,49 4,78 ± 0,17 340,48 ± 47,06 19,9 5,83 ± 0,65 3,93 ± 0,26 388,62 ± 36,79 26,6 5,33 ± 0,72 3,67 ± 0,15 327,64 ± 26,17
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A separate stage of root formation Rooting of Ortegocactus macdougallii shoots after 50 days. Means ± standard error. Eight vessels containing three explants each per treatment. IBA concentrations are given in µM in accordance with the methods and results descriptions. Data from Arellano-Perusquía et al. 2013, table 4. [Arellano-Perusquía et al. 2013, pp. 161, 164, Table 4] Medium Forming roots (%) Number of roots Root length (mm) ¼ MS, without regulators 58,34 ± 8,34 4,17 ± 0,48 11,32 ± 1,95 ½ MS, without regulators 37,49 ± 11,68 5,00 ± 1,37 9,96 ± 3,05 ½ MS + IBA 1,6 µM 45,83 ± 6,11 6,33 ± 0,77 8,99 ± 0,91 ½ MS + IBA 4,9 µM 75,03 ± 5,45 9,92 ± 1,91 8,37 ± 1,56
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What the habitat model represents Potential habitat area in three models. Values from table 2 of Solórzano et al. (2026). Variable set Potential area Model assessment in the publication Climate 34 469,25 km² Lowest AICc and highest TSS of the three models. Soil 77 082,23 km² Broadest projection, including beyond the confirmed range. Climate and soil 37 519,32 km² Model preferred by the authors for its agreement with field observations and highest pROC.
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Accepted names and the 2026 expansion Oehmea. Atlas classification and combinations from 2026. POWO status checked on 11 October 2026. Name accepted in the atlas Name in older accounts Context Oehmea beneckei Mammillaria beneckei Accepted by POWO. Buxbaum's 1951 combination. Oehmea sphacelata Mammillaria sphacelata Accepted by POWO. Guiggi's 2022 combination. Oehmea tonalensis Mammillaria tonalensis Accepted by POWO. Guiggi's 2022 combination. Mammillaria napina Mammillaria napina 2026 proposal. Oehmea napina . Mammillaria zephyranthoides Mammillaria zephyranthoides 2026 proposal. Oehmea zephyranthoides .
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Meristem size and stem size Meristem dome width in eight stages of the San Ángel material studied. Mean ± standard deviation, from table 2 of López-Ruiz et al. (2026). [López-Ruiz et al. 2026, p. 1333, Table 2; stage characteristics, pp. 1328, 1330–1334] Age after germination Dome width Stage characteristic 1 day 102,0 ± 30,2 µm Closed cotyledons, two recognisable layers. 3 days 94,2 ± 23,7 µm Tubercle primordia, three meristem layers. 7 days 140,2 ± 17,0 µm First visible spines. 15 days 128,1 ± 23,9 µm At least two tubercles with areoles. 30 days 137,0 ± 24,9 µm Differentiation of the areole-covered part of the stem. 90 days 212,1 ± 34,0 µm Meristem enlargement and spine stiffening. 1 year 224,7 ± 22,4 µm Distinct spine pattern. 4 years 771,7 ± 64,5 µm Flowering greenhouse plants.
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Different Mammillaria occupy different places Spatial associations in the study by Peters et al. (2008). “Positive” means more plants than expected in a given microhabitat, and “negative” means fewer. Accepted names follow POWO. Historical supertexta groups are retained separately. [Peters et al. 2008, p. 596, Table 1] [2-WWW] [2-WWW] Plant studied Shrubs Stones ≥20 cm M. dixanthocentron Association not significant in the test Positive M. hernandezii Negative Stones of this size were absent at the sites M. kraehenbuehlii Positive Positive M. supertexta (called M. lanata in the paper)Negative Negative M. napina Negative Association not significant in the test M. pectinifera Association not significant in the test Positive M. solisioides Negative Negative M. supertexta (under the same name in the paper)Negative Positive
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From a released seed to a seedling after the dry season Serotiny in Mammillaria pectinifera. Estimates for 238 m² in Texcala Quantity 1997–1998 1998–1999 Seeds released from retained fruits 35 508 32 259 Seeds released from ejected fruits 0 7634 All released seeds, estimate 35 508 39 893 Seedlings surviving the following dry season, estimate 17 95 Seedling establishment success, ‰ 0,47 2,38
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Medium, light and the importance of a specific stage Snowcap in vitro. Medium stages according to table 1 and the results of the 2023 paper. Stage Regulators in MS Sucrose Effect in the experiment Shoot maintenance, MSM-1 NAA 0,5 mg/l + KIN 0,3 mg/l 20 g/l Growth and material maintenance Offset stimulation, MSM-2 NAA 0,5 mg/l + KIN 0,3 mg/l + GA₃ 5 mg/l for 3 days 20 g/l Subsequent growth without GA₃ Rooting, MSM-3 NAA 0,5 mg/l + KIN 0,3 mg/l + ancymidol 5 mg/l 10 g/l Root development on offsets
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Flowers require different preparation Snowcap. Flower initiation in vitro after plant preparation and paclobutrazol treatment Paclobutrazol, mg/l Explants initiating flowers, % Flowers per explant 0 0 ± 0 0 ± 0 0,3 0 ± 0 0 ± 0 0,5 0 ± 0 0 ± 0 1 66 ± 6 1,40 ± 0,16 3 21 ± 5 0,37 ± 0,10 5 11 ± 4 0,24 ± 0,09
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Soil and landforms Selected site associations with soil and landscape map categories. Results of the frequency analysis in Aquino and co-authors (2021). [Aquino et al. 2021, pp. 4524–4525, Fig. 3] Species Categories with an excess of sites E. bokei Calcaric regosols E. pachyrhiza Calcaric xerosols and foothill zone E. pulchra Luvic kastanozems E. spinosior Luvic kastanozems and fold mountains E. polycephala Fold mountains
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Prism, druse, spherulite Selected biomineral forms and examples directly shown. Plant names follow the current atlas treatment. [De la Rosa-Tilapa et al. 2022, pp. 3–7, Fig. 2, 4–6] Form Tissue and example Observed characteristic Prisms Cortex of Epithelantha micromeris Crystals with distinct flat faces Druses Cortex of Kroenleinia grusonii (called Echinocactus grusonii in the paper) Clusters with projecting tips Crystal sand Cortex of Mammilloydia candida (called Mammillaria candida in the paper) Fine particles up to approximately 5 µm Spherulites Pith of Mammilloydia candida Radial arrangement and concentric bands Silica bodies Hypodermis of some Echinocactus and Ferocactus studied Amorphous deposits without birefringence
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Unusual polymers in former Escobaria Chemical identification of selected seed samples. The scope of the results corresponds to the material identifications in Chen and co-authors (2013). [Chen et al. 2013, pp. 203–208, Fig. 4–5] Current name Name used in the study Result Pelecyphora dasyacantha Escobaria dasyacantha Predominance of 5H. Almost homogeneous type in the NMR spectrum Pelecyphora lloydii Escobaria lloydii 5H and S units in chemical analysis products Pelecyphora zilziana Escobaria zilziana 5H and S, in proportions of approximately 63:37 in the analysed spectrum Pelecyphora sneedii subsp. sneedii Escobaria albicolumnaria Typical G/S lignin Pelecyphora vivipara Escobaria vivipara , arizonica and neomexicana materialTypical G/S lignin Epithelantha micromeris and E. greggii The same species names Lignin undetectable in the tests used
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Cell size and wall thickness Selected mean spine cell-wall thicknesses in the study by Řepka and Gebauer (2012). Measurements were made on hydrated preparations. Unit µm. [Řepka and Gebauer 2012, p. 141, Table 2] Material studied Epidermis Core fibres P. abdita (E. abdita )3,0 2,4 P. missouriensis , Montana sample (E. missouriensis subsp. missouriensis )4,5 3,7 P. missouriensis , Arizona sample (E. missouriensis subsp. navajoensis )3,6 4,0 P. minima (E. minima )3,6 3,3 P. hesteri (E. hesteri )4,2 3,8
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Seeds from different zones were transferred to common conditions Germination under the experimental conditions. Age means the estimated age of the tubercle zone, and the percentage is the mean reported in the publication. [Jiménez-Sierra et al. 2022, pp. 364–365, Tables 1–2] Material Estimated zone age Seeds / dishes Mean germination Fresh fruits current year's cohort 100 / 10 13% Zone 1 2,7 years 100 / 10 56% Zone 2 7,3 years 100 / 10 51% Zone 3 13,3 years 80 / 8 52% Zone 4 19,8 years 19 / 3 73% Zone 5 23,9 years 39 / 3 40%
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Self-pollen was often arrested in the style Response to hand self-pollination. Microscopy and fruit set were assessed in separate flowers. n denotes the size of each sample. [Martínez-Peralta et al. 2014b, p. 535, Table 1] Species under the current treatment Ovaries with pollen tubes (%) / n Flowers setting fruit (%) / n A. agavioides 25 / 28 6,9 / 29 A. bravoanus 33,3 / 3 0 / 5 A. fissuratus 7,7 / 39 1,5 / 67 A. kotschoubeyanus 18,9 / 53 19 / 124 A. retusus 3,8 / 26 4,3 / 69 A. scaphirostrus 0 / 37 2 / 50 A. trigonus 18,5 / 27 5,1 / 39
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Habitat loss and the emergence of new areas Change in area judged suitable by the BRT model. All values refer to the baseline area of habitats assessed as suitable. Net loss is the loss minus the newly suitable area. [Breslin et al. 2020, p. 13217, corrected Table 1] Horizon and RCP Loss of current area Newly suitable area Net loss 2050 · 2.6 46% 19% 27% 2050 · 4.5 48% 10% 38% 2050 · 6.0 47% 14% 33% 2050 · 8.5 52% 25% 27% 2070 · 2.6 50% 29% 21% 2070 · 4.5 50% 5% 45% 2070 · 6.0 50% 6% 44% 2070 · 8.5 55% 2% 53%
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Characters useful for identification Selected characters from the results and key of the 2019 revision. Areole dimensions were converted from centimetres to millimetres. Ranges apply to the material and treatment of this study. For E. pachyrhiza , former subsp. parvula was examined directly. [Aquino et al. 2019, pp. 606, 608, 610–612, 615] Species Areoles and spines Flower or underground part Epithelantha bokei 60–80 spines, persistent small projections on the spines Projections visible by electron microscopy E. cryptica Oval areoles, 0,60–0,75 mm wide Flowers 11–12 mm long E. greggii 19–30 spines, round areoles, 1,15–1,50 mm wide Ordinary slender roots E. ilariae Fewer than 40 spines, without persistent projections Flowers 17–18 mm long, opening in the evening E. micromeris Narrow areoles, 0,20–0,30 mm wide Flowers 6–7 mm long E. pachyrhiza White spines, elliptical areoles, 0,45–0,55 mm wide Thickened roots E. polycephala Brown spines, areoles 0,90–1,80 mm wide Thickened roots E. potosina 39–43 spines, areoles 0,50–0,70 mm long Thickened roots E. pulchra 40–55 spines, round areoles Thickened roots E. spinosior 19–25 spines, elliptical areoles Slender roots, part of the stem underground
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Self-pollen and foreign pollen in five treatments Fruit set in the studied Thelocactus leucacanthus population in Querétaro[Delgado-Ramírez et al. 2021, p. 798, Table 3] Treatment Flowers / plants Fruits Seeds per fruit, mean ± SD Freely accessible flowers 30 / 10 30, or 100% 65,76 ± 25,42 Hand pollination with foreign pollen 29 / 10 29, or 100% 61,52 ± 45,19 Hand self-pollination 24 / 8 0 No fruits formed Spontaneous self-pollination under cover 15 / 5 0 No fruits formed Apomixis trial 6 / 3 0 No fruits formed
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Nine species and different dominant plants Models of nine Thelocactus species. Data from Mosco's tables 1–2[Mosco 2019, p. 6, Tables 1–2] Species Sites Levins' niche breadth Highest permutation importance T. bicolor 213 0,282 Isothermality, 39,0% T. buekii 50 0,030 Precipitation of the wettest quarter, 36,6% T. conothelos 119 0,025 Precipitation of the driest quarter, 61,2% T. hastifer 11 0,178 Isothermality, 77,8% T. hexaedrophorus 195 0,099 Isothermality, 66,0% T. leucacanthus 73 0,050 Isothermality, 66,0% T. multicephalus 13 0,100 Isothermality, 33,0% T. rinconensis 47 0,062 Precipitation of the wettest quarter, 58,4% T. tulensis 117 0,020 Isothermality, 54,4%
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Two-year-old plants during summer warming Microclimate of the summer experiment. Means ± standard error[Aragón-Gastélum et al. 2017b, p. 95, microclimate results] Parameter Control plot Warming chambers Mean air temperature 21,5 ± 0,3°C 23,4 ± 0,4°C Mean daily maximum 33,5 ± 0,7°C 39,3 ± 1,0°C Mean daily minimum 12,5 ± 0,4°C 12,8 ± 0,4°C Relative air humidity 69,6 ± 2,1% 65,9 ± 2,4%
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Three Neoraimondia solution concentrations Water parameters after application of Neoraimondia arequipensis extract. Means ± SD[Choque-Quispe et al. 2020, p. 92, Table 1] Extract solution Turbidity removal Flocculation index pH BOD, mg O₂/l 1% 92,64 ± 0,33% 28,41 ± 0,94 6,75 ± 0,03 3,08 ± 0,02 2% 92,79 ± 0,18% 30,36 ± 0,11 6,93 ± 0,04 3,32 ± 0,02 3% 92,78 ± 0,09% 30,38 ± 0,11 7,02 ± 0,01 3,34 ± 0,03
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Structure helps identify the plant Examples of families and water-storage organs Family Example Storage organ Cactaceae Carnegiea gigantea Stem Aizoaceae Lithops ruschiorum Leaf Crassulaceae Crassula deceptor Leaf Asphodelaceae Aloe striata Leaf Asparagaceae Agave shawii Leaf Apocynaceae Hoodia gordonii Stem
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Data from studies and collections 26 results and parameters with measurement conditions ↗
Rootstocks Separate comparison of 19 grafting rootstocks ↗