Jiménez-Sierra C. L., Arroyo-Pérez E., Flores J., Zavala Hurtado J. A., Serotiny as a reproductive strategy in the specially protected species Ariocarpus kotschoubeyanus (Cactaceae), Plant Species Biology, 2022, 37(6):361-368, doi: 10.1111/1442-1984.12391
Observations of 30 adult Ariocarpus kotschoubeyanus document partial retention of mature seeds between tubercles. Tubercle formation rate estimates the age of five stem zones, and a germination test compares their material with fresh seeds at 25°C. The paper also describes germination on the mother plant and possible gradual release mechanisms for the bank.