Delgado-Ramírez V., Camacho-Velázquez A., Vázquez-Santana S., Biología reproductiva de Thelocactus leucacanthus ssp. schmollii (Cactaceae), un cactus microendémico de Querétaro, México, Botanical Sciences, 2021, 99(4):791-806, doi: 10.17129/botsci.2798
Flower anatomy, separation of male and female phases, five pollination trials and visitor observations in one population in Querétaro. Hand pollination with cross pollen and freely accessible flowers produced fruits. Self-pollination and apomixis trials produced no fruits.