Las Peñas M. L., Kiesling R., Bernardello G., Phylogenetic reconstruction of the genus Tephrocactus (Cactaceae) based on molecular, morphological, and cytogenetical data, Taxon, 2019, 68(4):714-730, doi: 10.1002/tax.12092
The phylogenetic analysis combines two chloroplast markers with morphology, chromosome counts, cytometry and rDNA mapping. It includes taxa treated as 12 Tephrocactus species in 2019. It documents substantial ploidy variation and reconstructs the history of selected traits.