Płachno B. J., Świątek P., Szymczak G., Can a stench be beautiful? – Osmophores in stem-succulent stapeliads (Apocynaceae-Asclepiadoideae-Ceropegieae-Stapeliinae), Flora, 2010, 205(2):101-105, doi: 10.1016/j.flora.2009.01.002
A microscopic study of osmophores in Orbea variegata and Boucerosia indica. It describes glandular papillae, subepidermal cells, organelles and routes of secretion release. It explains the anatomical basis for the intense scent of living flowers.