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@media(prefers-reduced-motion:reduce){*{scroll-behavior:auto!important;animation:none!important;transition:none!important}}
@media print{.header,footer,.article-toc,.related,.print-link,.skip,.diagram-switch{display:none!important}body{background:#fff;color:#000}.wrap{padding:0;max-width:none}.article-layout{display:block}.article-body{max-width:none}.article-heading h1{font-size:32pt}.article-section h2{font-size:19pt}.article-section p{font-size:11pt;line-height:1.5}.science-figure{break-inside:avoid;max-width:650px}.source-box{break-inside:avoid}.article-meta{padding:0 0 20px}.science-figure svg{max-height:240px}.article-heading{margin-bottom:25px}.article-section{margin-bottom:20px}}

.diagram-canvas{overflow-x:auto;max-width:100%;-webkit-overflow-scrolling:touch}.diagram-canvas svg{min-width:550px}.reading-card h3,.lead-feature h3,.news-teaser h3{font-family:var(--serif)}
.section-reference{font-size:.875rem!important;line-height:1.5!important;margin-top:20px!important}.section-reference a{color:#4c6a46;text-underline-offset:4px}.article-body table{font-size:1rem;line-height:1.65}.article-body caption{text-align:left;font-size:.875rem;font-weight:600;padding:0 0 12px}.article-toc{max-height:calc(100vh - 135px);overflow-y:auto}.article-toc button{font-size:.875rem;padding:12px 0}.article-meta,.reading-meta{font-size:.875rem}.science-figure .figure-label{font-size:.8125rem;line-height:1.5}.science-figure figcaption{font-size:.875rem;line-height:1.7}.science-figure .diagram-note{font-size:.9375rem;line-height:1.75}.diagram-switch button{font-size:.875rem}.article-sources a{font-size:.875rem}.article-sources .label{font-size:.75rem}.carbon-cycle,.organ-compare{display:grid;grid-template-columns:1fr 1fr;gap:18px;margin:22px 0}.carbon-column,.organ-compare>div{background:#fff;border:1px solid #d4dfcb;border-top:4px solid #829773;padding:22px}.carbon-column.cam{border-top-color:#234b3b}.carbon-label,.organ-type{font-size:.8125rem;font-weight:700;letter-spacing:.08em;color:#48613f}.carbon-subtitle{font-size:1rem;font-weight:600;margin:9px 0 20px;color:#263f30}.carbon-column ol{list-style:none;padding:0;margin:0;counter-reset:carbon}.carbon-column li{counter-increment:carbon;position:relative;padding:18px 0 18px 28px;border-top:1px solid #e0e7d9}.carbon-column li:before{content:counter(carbon);position:absolute;left:0;top:20px;font-size:.8rem;color:#718462}.carbon-column strong,.organ-compare strong{font-size:1rem;line-height:1.5;display:block;color:#263f30}.carbon-column li span{display:block;font-size:.875rem;line-height:1.7;margin-top:8px;color:#586451}.organ-compare strong{margin:12px 0}.organ-compare p,.organ-compare li{font-size:.9375rem!important;line-height:1.7!important}.organ-compare ol{padding-left:20px}.organ-compare li{margin-bottom:10px}.science-figure a{text-underline-offset:3px}.article-body h2{overflow-wrap:anywhere}.article-body a{overflow-wrap:anywhere}
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@media(max-width:600px){.hero{padding:0;align-items:center}.hero-copy{padding:45px 24px}.ticker{display:flex}.ticker span{display:block;margin:0!important}.ticker b{margin-bottom:5px}.diagram-canvas svg{min-width:550px}.diagram-canvas{padding-bottom:8px}}

/* Single-locus inheritance model */
.inheritance-model{padding:1.4rem;background:#f5f7ef;color:#244333}
.inheritance-model>p{text-align:center;font-size:1rem}
.inheritance-model .cross-parents{font:600 2rem Georgia,serif;margin:.2rem 0 1rem}
.inheritance-table{width:100%;max-width:33rem;margin:1rem auto;border-collapse:collapse;text-align:center}
.inheritance-table caption{font-size:.9rem;padding:.6rem;color:#465744}
.inheritance-table th,.inheritance-table td{border:1px solid #bdcbb1;padding:.8rem;text-align:center!important}
.inheritance-table thead th,.inheritance-table tbody th{background:#e2ebd8}
.inheritance-table td{font:600 1.5rem Georgia,serif;background:#fff}
.inheritance-result{display:flex;justify-content:center;gap:1rem;flex-wrap:wrap;margin:1rem 0}
.inheritance-result span{display:flex;flex-direction:column;align-items:center;gap:.2rem;min-width:5rem;font-size:1.1rem}
.inheritance-result strong{font-size:1.25rem}
.subfamily-grid{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:1px;background:var(--line);border:1px solid var(--line);margin:32px 0 52px}.growth-note{border-top:1px solid var(--line);padding-top:10px;margin-top:14px}.subfamily-grid .growth p{font-size:.95rem}
@media(max-width:850px){.subfamily-grid{grid-template-columns:repeat(2,minmax(0,1fr))}}
@media(max-width:600px){.subfamily-grid{grid-template-columns:1fr}}

/* Aligned, selectable reference tables and independent scientific plates. */
.reference-table-wrap{max-width:100%;overflow-x:auto;margin:28px 0 24px;border:1px solid #cbd7c4;border-radius:4px;background:#fff;scrollbar-color:#718566 #edf2e8}
.reference-table-wrap:focus-visible{outline:3px solid #648541;outline-offset:4px}
.article-body .reference-table{table-layout:fixed;min-width:660px;width:100%;font-size:1rem;line-height:1.65}
.article-body .reference-table caption{padding:18px 20px;background:#f0f4e9;color:#284737;font-size:1rem;border-bottom:1px solid #cbd7c4}
.reference-table th,.reference-table td{text-align:left;vertical-align:top;padding:16px 18px;overflow-wrap:anywhere;min-width:0}
.reference-table thead th{font-size:.875rem;background:#234b3b;color:#fff;letter-spacing:.015em}
.reference-table thead th{width:auto}
/* The five-column stock table uses explicit colgroup widths. */
.rootstock-table{table-layout:fixed;min-width:1050px;width:100%}.rootstock-table th:nth-child(n){width:auto}.rootstock-table .stock-name{width:18%}.rootstock-table .stock-original{width:18%}.rootstock-table .stock-use{width:25%}.rootstock-table .stock-limits{width:25%}.rootstock-table .stock-source{width:14%}.rootstock-table th,.rootstock-table td{word-break:normal;overflow-wrap:break-word;hyphens:auto}.rootstock-table thead th{white-space:normal}.rootstock-table .lit-ref{white-space:normal}

.reference-table tbody th{font-size:1rem;font-weight:600;background:transparent;color:#244b38}
.reference-table tbody tr:nth-child(even){background:#f5f8f0}.reference-table tbody tr:hover{background:#eaf1e1}
.reference-table tbody tr:last-child>*{border-bottom:0}
.reference-table i{font-family:Georgia,'Times New Roman',serif;font-size:1.06em}
.method-grid{display:grid;grid-template-columns:repeat(auto-fit,minmax(250px,1fr));gap:18px;margin:24px 0}
.method-panel{min-width:0;padding:18px;background:#fff;border:1px solid #cfdbc7;border-radius:4px}
.method-panel h3{font-family:var(--serif);font-size:1.35rem;line-height:1.3;margin:0 0 16px}
.method-panel svg{display:block;width:100%;height:auto;max-width:340px;margin:0 auto 16px}
.article-section .method-panel p{font-size:1rem;line-height:1.75;margin:0}
.method-figure .figure-label{font-size:.875rem;letter-spacing:.055em}
@media(max-width:600px){.method-grid{grid-template-columns:minmax(0,1fr)}.method-panel{padding:16px}.method-panel svg{min-width:0}.article-body .reference-table{min-width:660px}.reference-table-wrap{margin-right:0;margin-left:0}.method-panel h3{font-size:1.4rem}}
@media print{.reference-table-wrap{overflow:visible;border:0}.article-body .reference-table{min-width:0;font-size:9pt;table-layout:fixed}.reference-table th,.reference-table td{padding:8px}.reference-table thead{display:table-header-group}.reference-table tr{break-inside:avoid}.method-figure{break-inside:auto}.method-panel{break-inside:avoid}.method-grid{grid-template-columns:1fr 1fr}}

/* Collection directories keep the existing typography and color system. */
.collection-entry{padding-top:48px;padding-bottom:48px}.diagram-directory{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:16px;margin:25px 0 55px}.diagram-tile{display:block;border:1px solid var(--line,#d8ded1);border-radius:12px;padding:25px;background:#f7f8f1;color:#163d31;text-decoration:none}.diagram-tile:hover{background:#e8efdc}.diagram-tile h3{font-size:1.15rem;line-height:1.45;text-transform:none;margin:18px 0}.diagram-tile span:last-child{font-size:.85rem}.archive-fact{padding:18px 0;border-bottom:1px solid #dbe1d3}.archive-fact h3{margin-bottom:12px}.evidence-note{font-size:.94rem;color:#596958;border-left:3px solid #b2bf9e;padding-left:16px}.procedure-steps{padding-left:1.5em}.procedure-steps li{padding:0 0 20px 12px;line-height:1.8}.procedure-steps li::marker{font-weight:700;color:#36573f}.table-section{padding:28px 0;scroll-margin-top:100px}.index-pagination{display:flex;gap:12px;margin:24px 0}.index-pagination button:disabled{opacity:.4;cursor:default}.reference-table th,.reference-table td{vertical-align:top}.hub-links button{text-align:left}.diagram-reading{margin:24px 0}.reference-table a{overflow-wrap:anywhere}#name-results{scroll-margin-top:110px}nav a{white-space:nowrap}@media(min-width:901px){.header{gap:20px}nav{gap:17px}nav a{font-size:.82rem}}@media(max-width:900px){.diagram-directory{grid-template-columns:repeat(2,minmax(0,1fr))}}@media(max-width:560px){.diagram-directory{grid-template-columns:1fr}.collection-entry{padding-top:32px}.diagram-tile{padding:20px}}

.reference-table .numeric{font-variant-numeric:tabular-nums;white-space:nowrap;text-align:right}

/* Editorial headings have a calmer scale and balanced line breaks. */
.article-heading h1,.page-head h1{font-family:var(--serif);font-size:clamp(2rem,4.2vw,3.55rem);font-weight:400;line-height:1.16;letter-spacing:-.025em;text-wrap:balance;max-width:24ch;overflow-wrap:normal;hyphens:auto}.article-section h2,.article-body h2,.content h2{font-size:clamp(1.55rem,2.5vw,2.15rem);font-weight:400;line-height:1.25;letter-spacing:-.015em;text-wrap:balance;overflow-wrap:normal;hyphens:auto}.reading-card h3,.diagram-tile h3{font-weight:400;line-height:1.3;text-wrap:balance}.article-heading .article-deck{max-width:70ch}@media(max-width:600px){.article-heading h1,.page-head h1{font-size:2.1rem;line-height:1.18;max-width:100%}}

.botanical-board{max-width:850px;margin:32px auto 40px}.botanical-board img{width:100%;height:auto;border-radius:5px}.botanical-board figcaption{font-size:.9rem;line-height:1.65;color:var(--muted);margin-top:12px}.botanical-gallery{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:32px;margin:40px 0}.botanical-gallery figure{margin:0}.botanical-gallery img{width:100%;height:auto;display:block;border-radius:5px}.botanical-gallery h2{font-size:1.65rem;line-height:1.25;margin:20px 0 10px}.botanical-gallery p{font-size:1rem;line-height:1.75}.gallery-promo{margin:32px 0;padding:20px;background:#eee9db;border-radius:8px;max-width:860px}.gallery-promo>a{display:flex;gap:28px;align-items:center;text-decoration:none}.gallery-promo img{width:180px;height:180px;object-fit:contain}.gallery-promo strong{display:block;font-family:var(--serif);font-size:1.9rem;font-weight:400;line-height:1.25;margin:10px 0}.gallery-promo span span:last-child{font-size:1rem}.taxonomy-counts{display:flex;flex-wrap:wrap;gap:20px 35px;margin:28px 0}.taxonomy-counts span{font-size:.9rem}.taxonomy-counts b{display:block;font-size:1.8rem;font-weight:400}.full-taxonomy-tree{border-top:1px solid var(--line)}.taxon-genus{border-bottom:1px solid var(--line);padding:0 10px}.taxon-genus>summary{padding:20px 8px;cursor:pointer;font-size:1.3rem}.taxon-genus>summary small{font-size:.875rem;color:var(--muted);margin-left:20px}.genus-content{padding:0 10px 16px 22px}.taxon-list{list-style:none;padding:0;margin:0}.taxon-row{border-top:1px solid #dde3d5}.taxon-row summary{cursor:pointer;padding:15px 0;line-height:1.65}.taxon-row summary small{font-size:.9rem;color:var(--muted)}.taxon-tags{display:inline-flex;gap:5px;margin-left:12px}.taxon-tags .tag{font-size:.75rem}.taxon-info{padding:10px 16px 25px;border-left:2px solid #b6c6a1}.taxon-info h4{font-size:1rem;margin:20px 0 12px}.synonym-list{list-style:none;padding:0;columns:2;column-gap:35px}.synonym-list li{break-inside:avoid;margin-bottom:12px;font-size:.95rem;line-height:1.6}.synonym-list small{display:block;color:var(--muted);font-size:.8rem}.higher-genera{display:flex;gap:10px;flex-wrap:wrap;padding:20px}.name-status{margin:30px 0}.name-status li{margin-bottom:10px}.name-status h3{font-size:1.5rem}.full-taxonomy-tree details[open]>summary{color:#194d3c}.full-taxonomy-tree summary:focus-visible{outline:2px solid #194d3c;outline-offset:4px}@media(max-width:850px){.botanical-gallery{grid-template-columns:repeat(2,minmax(0,1fr))}.synonym-list{columns:1}}@media(max-width:560px){.botanical-gallery{grid-template-columns:1fr;gap:25px}.gallery-promo>a{display:block}.gallery-promo img{width:100%;height:auto;max-height:320px}.taxon-genus>summary small{display:block;margin-left:20px}.taxon-tags{display:flex;margin:8px 0 0}.genus-content{padding-left:10px}.taxon-info{padding-left:12px}.taxonomy-counts{gap:15px}.taxonomy-counts span{flex-basis:44%}}

@media print{.rootstock-table{min-width:0;font-size:8pt}.rootstock-table th,.rootstock-table td{padding:6px}}

/* Directory previews and unobtrusive topic icons for texts without figures. */
.diagram-tile-preview{display:flex;align-items:center;justify-content:center;height:128px;margin:0 0 1rem;overflow:hidden;background:#fff;border-radius:8px}
.diagram-tile-preview img,.diagram-tile-preview svg{display:block;width:100%;height:100%;object-fit:contain;pointer-events:none}
.diagram-tile-preview svg{max-width:100%;color:var(--green)}
.topic-preview{color:#526e4a;background:#edf1e5;padding:18px}
.topic-preview svg{width:100%;height:100%}
.article-topic-icon{color:#526e4a;background:#edf1e5;border:1px solid #d4dfc9;border-radius:18px;padding:20px;width:112px;height:112px;flex:0 0 112px;align-self:center}
.article-topic-icon svg{width:100%;height:100%}
@media(max-width:680px){.article-topic-icon{width:76px;height:76px;padding:12px;flex-basis:76px;align-self:flex-start}.diagram-tile-preview{height:110px}}